Mirabilis laevis |
Mirabilis glabra |
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desert four o'clock, desert wishbone, desert wishbone-bush, wishbone bush |
smooth four-o'clock |
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Stems | decumbent to erect, few and clambering through other vegetation to many, and then usually forming densely leafy and compact clumps, 1.5–15 dm, herbaceous, suffrutescent, or woody basally, glabrous, scabrous, puberulent, or villous, often glandular. |
erect or ascending, sparsely to moderately leafy mostly in proximal 2/3, narrowly forked, usually with main axis, 5–20 dm, glabrous, glandular-puberulent, or puberulent with curved hairs, when pubescent, hairs usually densest distally. |
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Leaves | spreading; petiole 0.1–2.2 cm; blade ovate, deltate-ovate, ovate-rhombic, subreniform, 1–4(–5.5) × 0.5–3.5(–5) cm, fleshy to slightly succulent, base cordate, truncate, or broadly obtuse, apex acute, obtuse, or rounded, surfaces glabrous, scabrous, puberulent, or villous, often glandular. |
strongly ascending at 5–25°, progressively reduced toward inflorescence; petiole 0–1.3 cm; blade glaucous or grayish green, linear to narrowly ovate or ovate-oblong, 5–10 × 0.2–7.5 cm, thick and somewhat coriaceous, base attenuate to obtuse, round, truncate, or subcordate, apex acute to obtuse, occasionally round, surfaces glabrous to short pilose. |
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Inflorescences | widely cymose, or ± thyrsoid, involucres clustered, and nearly sessile at ends of branches, or solitary in axils on peduncles 3–12 mm; involucres 3–7 mm, lobes narrowly to broadly triangular, or triangular-lanceolate, base 30–50% of height. |
usually terminal, sometimes also with short branches in distal axils, open, with main axis and short branches; peduncle 5–10 mm, usually spreading glandular-pilose, sometimes merely puberulent or glabrous, crosswalls of hairs pale; involucres pale green, widely bell-shaped, 3–6 mm in flower, 7–12 mm in fruit, glabrous except for few small hairs on margins to spreading glandular-pilose, 60–90% connate, lobes broadly ovate, rounded. |
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Flowers | 1(–2) per involucre; perianth white, pink, or shades of purple, 1–1.6 cm. |
1–3 per involucre; perianth white to pale pink, 0.6–0.9 cm. |
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Fruits | gray, dark brown, or nearly black, often mottled with dark brown or black, with or without 10 pale, diffuse lines, ovoid, obovoid, or nearly spheric, 3–5.5 mm, smooth or moderately rugose. |
grayish to greenish brown (ribs usually paler), narrowly obovate and tapering at both ends, 4–5.5 mm, usually glabrous, sometimes very lightly puberulent with hairs 0.1 mm; ribs narrow and keel-like to acute or acute-rounded, 0.2–0.5 times width of sulci, 0.5 times as wide as high, smooth; sulci lightly to prominently tuberculate. |
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Mirabilis laevis |
Mirabilis glabra |
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Phenology | Flowering late spring–early fall. | |||||||||
Habitat | Sandy soils in grasslands or among oaks, juniper, or mesquite, disturbed areas | |||||||||
Elevation | 500-2100 m (1600-6900 ft) | |||||||||
Distribution |
AZ; CA; NV; OR; UT; nw Mexico
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AZ; CO; KS; NE; NM; OK; TX; UT; Mexico (Chihuahua)
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Discussion | Varieties 4 (3 in the flora). Mirabilis laevis is a complex of poorly differentiated forms that differ to a greater or lesser extent primarily by perianth color, pubescence, and habit, characteristics that show imperfect geographic consistency. In general, white-flowered plants occur in arid areas east of the southern California mountains, and magenta-flowered plants occur west of the mountains; in the arid regions viscid-pubescent plants occur to the south, less viscid plants to the north. Sympatry and intergradation are frequent in the southern Sierra Nevada, southward along the east side of the southern California mountains, and on the northern portion of the peninsula of Baja California. The variety laevis, which is glabrous or glabrate, is restricted to the immediate coast and islands in the vicinity of Bahía Magdalena in Baja California Sur. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
As traditionally treated, Mirabilis carletonii (stems short pilose) and M. exaltata (stems glabrous), both from the eastern portion of the range of M. glabra, have leaves broader than 10 mm. Correlated with this is the presence of two to three flowers in the involucres. The narrower-leaved M. glabra, in the strict sense, has one or occasionally two flowers per involucre. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 4, p. 47. | FNA vol. 4, p. 56. | ||||||||
Parent taxa | Nyctaginaceae > Mirabilis > sect. Oxybaphoides | Nyctaginaceae > Mirabilis > sect. Oxybaphus | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Oxybaphus laevis, Hesperonia laevis | Oxybaphus glaber, Allionia carletonii, Allionia exaltata, Allionia glabra, M. carletonii, M. exaltata, Oxybaphus carletonii, Oxybaphus exaltatus | ||||||||
Name authority | (Bentham) Curran: Proc. Calif. Acad. Sci., ser. 2, 1: 235. (1888) | (S. Watson) Standley: Publ. Field Mus. Nat. Hist., Bot. Ser. 8: 304. (1931) | ||||||||
Web links |