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Ross' sandwort

Howell's sandwort, Howell's stitchwort

Habit Plants perennial, densely pulvinate to loosely cespitose. Plants annual.
Taproots

stout, woody.

moderately stout.

Stems

ascending to spreading, green or often purple, 1–3 cm, glabrous, internodes of flowering stems 0.2–1 times as long as leaves.

erect to spreading, green, becoming purple, 12–30 cm, mostly glabrous, internodes of all stems 2–7 times as long as leaves.

Leaves

overlapping, ± tightly (vegetative), ± evenly spaced proximally (cauline), connate-perfoliate proximally, with tight, herbaceous sheath 0.2–0.3 mm;

blade upwardly curved, green or often purple, keeled, prominently 1-veined abaxially, subulate, 3-angled, 1–4 × 0.5–0.7 mm, flexuous, margins rounded, herbaceous, smooth, apex green to purple, rounded, navicular, shiny, glabrous;

axillary leaves well developed.

sometimes overlapping proximally, often connate proximally, with loose, scarious to herbaceous sheath 0.2–0.5 mm;

blade straight to recurved, green, becoming purple, concave, prominently 1(–3)-veined abaxially, linear-lanceolate, 5–15 × 1–2.5 mm, rigid, margins not thickened, slightly scarious, smooth, apex green to purple, obtuse to acute, often apiculate, flat to navicular, dull, densely stipitate-glandular;

axillary leaves absent.

Inflorescences

solitary flowers, axillary or terminal (rarely present);

bracts absent.

5–25+-flowered, open cymes;

bracts lanceolate to ovate, herbaceous.

Pedicels

0.1–2 cm, glabrous.

0.3–4+ cm, stipitate-glandular.

Flowers

hypanthium disc-shaped;

sepals 1-veined, oblong-ovate (herbaceous portion usually purple, oblong-ovate), 1.5–2.5 mm, not enlarging in fruit, apex often purple, obtuse to acuminate, navicular, not hooded, glabrous;

petals obovate to spatulate, 1.5–2 times as long as sepals, apex obtuse, entire.

hypanthium disc-shaped;

sepals weakly veined proximally, ovate (herbaceous portion ovate to narrowly so), 1.9–3 mm, not enlarging in fruit, apex often purple, ± acute to acuminate, not hooded, stipitate-glandular proximally;

petals oblanceolate, 1.8–2.3 times as long as sepals, apex rounded, entire.

Capsules

on stipe ca. 0.1–0.2 mm, spheric, 1.5–2.5 mm, equaling sepals.

on stipe ca. 0.2 mm, broadly ovoid, 2.7–3.5 mm, usually longer than sepals.

Seeds

brown, suborbiculate, compression unknown, ca. 0.6 mm, obscurely reticulate.

blackish brown, suborbiculate with radicle prolonged to rounded beak, somewhat compressed, 1.4–1.7 mm, tuberculate;

tubercles low, rounded, often elongate.

2n

= 58 (Russia), 60.

Minuartia rossii

Minuartia howellii

Phenology Flowering spring–summer. Flowering spring–summer.
Habitat Wet, turfy, gravelly, or sandy calcareous barrens, high arctic, alpine tundra, heathlands Chaparral, Jeffrey pine-oak woodlands, serpentine
Elevation 0-500 m (0-1600 ft) 500-1000 m (1600-3300 ft)
Distribution
from FNA
AK; NT; NU; YT; Greenland; Europe (Spitzbergen); Asia (Russian Far East)
[BONAP county map]
from FNA
CA; OR
[BONAP county map]
Discussion

Minuartia rossii is the northernmost member of the M. rossii complex (S. J. Wolf et al. 1979; B. Maguire 1958), a pulvinate species of moist arctic areas. While specimens occasionally have many flowers, some specimens have few if any, instead reproducing via small axillary fascicles of leaves or short shoots in the upper leaf axils (see also Ö. Nilsson 2001).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 133. FNA vol. 5, p. 127.
Parent taxa Caryophyllaceae > subfam. Alsinoideae > Minuartia Caryophyllaceae > subfam. Alsinoideae > Minuartia
Sibling taxa
M. arctica, M. austromontana, M. biflora, M. californica, M. caroliniana, M. cismontana, M. cumberlandensis, M. dawsonensis, M. decumbens, M. douglasii, M. drummondii, M. elegans, M. glabra, M. godfreyi, M. groenlandica, M. howellii, M. macrantha, M. macrocarpa, M. marcescens, M. michauxii, M. muscorum, M. nuttallii, M. obtusiloba, M. patula, M. pusilla, M. rosei, M. rubella, M. stolonifera, M. stricta, M. tenella, M. uniflora, M. yukonensis
M. arctica, M. austromontana, M. biflora, M. californica, M. caroliniana, M. cismontana, M. cumberlandensis, M. dawsonensis, M. decumbens, M. douglasii, M. drummondii, M. elegans, M. glabra, M. godfreyi, M. groenlandica, M. macrantha, M. macrocarpa, M. marcescens, M. michauxii, M. muscorum, M. nuttallii, M. obtusiloba, M. patula, M. pusilla, M. rosei, M. rossii, M. rubella, M. stolonifera, M. stricta, M. tenella, M. uniflora, M. yukonensis
Synonyms Arenaria rossii, Alsinanthe rossii, Alsinopsis rossii, Arenaria rossii var. apetala, M. orthotrichoides, M. rolfii, M. rossii var. orthotrichoides Arenaria howellii, Alsinopsis howellii
Name authority (R. Brown ex Richardson) Graebner: in P. F. A. Ascherson et al., Syn. Mitteleur. Fl. 5(1): 772. (1918) (S. Watson) Mattfeld: Bot. Jahrb. Syst. 57(Beibl. 126): 27. (1921)
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