Minuartia patula
Minuartia rubella
Pitcher's stitchwort
arctic sandwort, beautiful sandwort, boreal sandplant, boreal stitchwort, minuartie rougeâtre, red seed sandwort, reddish sandwort
filiform.
filiform to somewhat thickened;
rhizomes absent.
erect to ascending, green, 5–30 cm, glabrous or sometimes stipitate-glandular distally or throughout, internodes of all stems 1–7 times as long as leaves; wintering stems absent.
ascending to erect, green, 2–8(–18) cm, moderately to densely stipitate-glandular (very rarely glabrous), internodes of stems 1–10 times as long as leaves; trailing stems absent.
overlapping proximally, connate proximally, with loose, scarious to herbaceous sheath 0.1–0.5 mm;
blade straight to variously curved, green, flat, prominently 1-veined abaxially, linear, 2–20 × 0.5–1.5(–1.8) mm, flexuous, margins not thickened, herbaceous, smooth, apex green or purple, blunt to acute, flat, ± shiny, glabrous to stipitate-glandular;
axillary leaves absent.
overlapping, ± tightly, distally (cauline), concentrated proximally (cauline), connate proximally, with often loose, usually scarious sheath 0.2–0.7 mm;
blade ± straight or outwardly curved, green, flat to 3-angled, prominently 3-veined abaxially, subulate, 1.5–10 × 0.3–1.3 mm, flexuous, margins not thickened, scarious, smooth, apex green or purple, acute to apiculate, often navicular, shiny, sparsely to densely ciliate, often stipitate-glandular;
axillary leaves present among vegetative leaves.
5–30-flowered, open cymes;
bracts subulate to ovate, herbaceous.
3–7+-flowered, open cymes or rarely flower solitary, terminal;
bracts broadly subulate to narrowly lanceolate, herbaceous, margins scarious.
0.3–3 cm, stipitate-glandular.
0.2–1.5 cm, densely stipitate-glandular.
hypanthium shallowly disc-shaped;
sepals prominently (3- or) 5-veined, narrowly to broadly lanceolate (herbaceous portion narrowly to broadly lanceolate), 4–5.5 mm, not enlarging in fruit, apex green or purple, narrowly acute to acuminate, not hooded, glabrous to sparsely stipitate-glandular;
petals obovate, 1.5–2.2(–3) times as long as sepals, apex rounded, broadly notched.
hypanthium disc-shaped;
sepals prominently 3-veined, ovate to lanceolate (herbaceous portion oblong to narrowly ovate), 2.5–3.2 mm, not enlarging in fruit, apex green to purple, acute to acuminate, not hooded, stipitate-glandular;
petals elliptic, 0.8–1.3 times as long as sepals, apex rounded, entire.
on stipe ca. 0.1 mm or shorter, narrowly ellipsoid, 3–4.2 mm, shorter than sepals.
on stipe ca. 0.2 mm, ovoid, 4.5–5 mm, longer than sepals.
reddish brown to black, suborbiculate, radicle obscure, slightly compressed, 0.5–0.6 mm, tuberculate;
tubercles low, rounded.
reddish brown, suborbiculate with radicle prolonged into beak, somewhat compressed, 0.4–0.5 mm, tuberculate;
tubercles low, elongate, rounded (to angled on edge) (50x).
= 24.
Minuartia patula
Minuartia rubella
Minuartia patula and the related M. muscorum have received little attention in comparison to the granite-outcrop minuartias, the M. uniflora complex. J. A. Steyermark (1941) studied these taxa and described three forms, based chiefly on pubescence variation. Plants entirely glabrous [forma pitcheri (Nuttall) Steyermark] and those with sepals and pedicels somewhat stipitate-glandular (forma media Steyermark) were segregated from densely stipitate-glandular plants (forma patula). We do not feel that such variations deserve formal taxonomic recognition. Forma robusta, as defined by Steyermark, is here referred to M. muscorum.
Most specimens of Minuartia patula have prominently five-veined sepals (seen especially easily in the glabrous forms); occasional plants from Georgia, Kentucky, and Virginia have glabrous sepals with only three strong veins, resembling those of M. muscorum; in other features, including the seeds, they are clearly referable to M. patula. The status of the plants with three-veined sepals remains ambiguous; J. A. Steyermark (1941) included them in his forma media and B. Maguire (1951) included them (in our opinion incorrectly) in his var. robusta.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Distinct among the arctic/alpine Minuartia species with its stiff, three-veined leaves, M. rubella is a circumpolar calciphile. We follow Ö. Nilsson (2001) in not recognizing infraspecific taxa that have been described based at least partly on pubescence. Variety propinqua has been applied to glabrous plants, which occur infrequently and sporadically throughout the range of the species. Where they do occur they are often intermixed with sparsely stipitate-glandular plants. This glabrous variety is rarely encountered in western North America.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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