Minuartia patula |
Minuartia rossii |
|
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Pitcher's stitchwort |
Ross' sandwort |
|
Habit | Plants winter annual or annual. | Plants perennial, densely pulvinate to loosely cespitose. |
Taproots | filiform. |
stout, woody. |
Stems | erect to ascending, green, 5–30 cm, glabrous or sometimes stipitate-glandular distally or throughout, internodes of all stems 1–7 times as long as leaves; wintering stems absent. |
ascending to spreading, green or often purple, 1–3 cm, glabrous, internodes of flowering stems 0.2–1 times as long as leaves. |
Leaves | overlapping proximally, connate proximally, with loose, scarious to herbaceous sheath 0.1–0.5 mm; blade straight to variously curved, green, flat, prominently 1-veined abaxially, linear, 2–20 × 0.5–1.5(–1.8) mm, flexuous, margins not thickened, herbaceous, smooth, apex green or purple, blunt to acute, flat, ± shiny, glabrous to stipitate-glandular; axillary leaves absent. |
overlapping, ± tightly (vegetative), ± evenly spaced proximally (cauline), connate-perfoliate proximally, with tight, herbaceous sheath 0.2–0.3 mm; blade upwardly curved, green or often purple, keeled, prominently 1-veined abaxially, subulate, 3-angled, 1–4 × 0.5–0.7 mm, flexuous, margins rounded, herbaceous, smooth, apex green to purple, rounded, navicular, shiny, glabrous; axillary leaves well developed. |
Inflorescences | 5–30-flowered, open cymes; bracts subulate to ovate, herbaceous. |
solitary flowers, axillary or terminal (rarely present); bracts absent. |
Pedicels | 0.3–3 cm, stipitate-glandular. |
0.1–2 cm, glabrous. |
Flowers | hypanthium shallowly disc-shaped; sepals prominently (3- or) 5-veined, narrowly to broadly lanceolate (herbaceous portion narrowly to broadly lanceolate), 4–5.5 mm, not enlarging in fruit, apex green or purple, narrowly acute to acuminate, not hooded, glabrous to sparsely stipitate-glandular; petals obovate, 1.5–2.2(–3) times as long as sepals, apex rounded, broadly notched. |
hypanthium disc-shaped; sepals 1-veined, oblong-ovate (herbaceous portion usually purple, oblong-ovate), 1.5–2.5 mm, not enlarging in fruit, apex often purple, obtuse to acuminate, navicular, not hooded, glabrous; petals obovate to spatulate, 1.5–2 times as long as sepals, apex obtuse, entire. |
Capsules | on stipe ca. 0.1 mm or shorter, narrowly ellipsoid, 3–4.2 mm, shorter than sepals. |
on stipe ca. 0.1–0.2 mm, spheric, 1.5–2.5 mm, equaling sepals. |
Seeds | reddish brown to black, suborbiculate, radicle obscure, slightly compressed, 0.5–0.6 mm, tuberculate; tubercles low, rounded. |
brown, suborbiculate, compression unknown, ca. 0.6 mm, obscurely reticulate. |
2n | = 58 (Russia), 60. |
|
Minuartia patula |
Minuartia rossii |
|
Phenology | Flowering spring–early summer. | Flowering spring–summer. |
Habitat | Prairies, meadows, limestone barrens, and rocky outcrops in sandy, clayey, or gravelly soils | Wet, turfy, gravelly, or sandy calcareous barrens, high arctic, alpine tundra, heathlands |
Elevation | 0-500 | 0-500 m (0-1600 ft) |
Distribution |
AL; AR; GA; IL; IN; KS; KY; LA; MO; MS; OH; OK; PA; TN; TX; VA; WI |
AK; NT; NU; YT; Greenland; Europe (Spitzbergen); Asia (Russian Far East) |
Discussion | Minuartia patula and the related M. muscorum have received little attention in comparison to the granite-outcrop minuartias, the M. uniflora complex. J. A. Steyermark (1941) studied these taxa and described three forms, based chiefly on pubescence variation. Plants entirely glabrous [forma pitcheri (Nuttall) Steyermark] and those with sepals and pedicels somewhat stipitate-glandular (forma media Steyermark) were segregated from densely stipitate-glandular plants (forma patula). We do not feel that such variations deserve formal taxonomic recognition. Forma robusta, as defined by Steyermark, is here referred to M. muscorum. Most specimens of Minuartia patula have prominently five-veined sepals (seen especially easily in the glabrous forms); occasional plants from Georgia, Kentucky, and Virginia have glabrous sepals with only three strong veins, resembling those of M. muscorum; in other features, including the seeds, they are clearly referable to M. patula. The status of the plants with three-veined sepals remains ambiguous; J. A. Steyermark (1941) included them in his forma media and B. Maguire (1951) included them (in our opinion incorrectly) in his var. robusta. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Minuartia rossii is the northernmost member of the M. rossii complex (S. J. Wolf et al. 1979; B. Maguire 1958), a pulvinate species of moist arctic areas. While specimens occasionally have many flowers, some specimens have few if any, instead reproducing via small axillary fascicles of leaves or short shoots in the upper leaf axils (see also Ö. Nilsson 2001). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 5, p. 131. | FNA vol. 5, p. 133. |
Parent taxa | Caryophyllaceae > subfam. Alsinoideae > Minuartia | Caryophyllaceae > subfam. Alsinoideae > Minuartia |
Sibling taxa | ||
Synonyms | Arenaria patula, Alsinopsis patula, Alsinopsis pitcheri, Sabulina patula | Arenaria rossii, Alsinanthe rossii, Alsinopsis rossii, Arenaria rossii var. apetala, M. orthotrichoides, M. rolfii, M. rossii var. orthotrichoides |
Name authority | (Michaux) Mattfeld: Bot. Jahrb. Syst. 57(Beibl. 126): 28. (1921) | (R. Brown ex Richardson) Graebner: in P. F. A. Ascherson et al., Syn. Mitteleur. Fl. 5(1): 772. (1918) |
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