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Pitcher's stitchwort

cismontane minuartia

Habit Plants winter annual or annual. Plants annual.
Taproots

filiform.

filiform.

Stems

erect to ascending, green, 5–30 cm, glabrous or sometimes stipitate-glandular distally or throughout, internodes of all stems 1–7 times as long as leaves; wintering stems absent.

erect, green or reddish purple, (5–)8–20(–25) cm, glabrous, internodes of all stems 5–7 times as long as leaves.

Leaves

overlapping proximally, connate proximally, with loose, scarious to herbaceous sheath 0.1–0.5 mm;

blade straight to variously curved, green, flat, prominently 1-veined abaxially, linear, 2–20 × 0.5–1.5(–1.8) mm, flexuous, margins not thickened, herbaceous, smooth, apex green or purple, blunt to acute, flat, ± shiny, glabrous to stipitate-glandular;

axillary leaves absent.

not overlapping, connate proximally, with loose, scarious sheath 0.1–0.3 mm;

blade green or reddish purple, 3-veined proximally, midvein prominent abaxially, lateral veins 0.2–0.25 times as long as blade, straight to outwardly curved, flat, lance-attenuate to linear, 2–7(–9) × 0.5–1.2(–1.8) mm, flexuous, margins not thickened, scarious proximally, smooth, apex green or purple, rounded to acute, often mucronate, shiny, glabrous;

axillary leaves occasionally present.

Inflorescences

5–30-flowered, open cymes;

bracts subulate to ovate, herbaceous.

5–20-flowered, open cymes;

bracts subulate, herbaceous, often scarious-margined proximally.

Pedicels

0.3–3 cm, stipitate-glandular.

(0.7–)1–3(–3.5) cm, glabrous.

Flowers

hypanthium shallowly disc-shaped;

sepals prominently (3- or) 5-veined, narrowly to broadly lanceolate (herbaceous portion narrowly to broadly lanceolate), 4–5.5 mm, not enlarging in fruit, apex green or purple, narrowly acute to acuminate, not hooded, glabrous to sparsely stipitate-glandular;

petals obovate, 1.5–2.2(–3) times as long as sepals, apex rounded, broadly notched.

hypanthium disc-shaped;

sepals strongly 3(–5)-veined, lance-linear to lanceolate (herbaceous portion narrowly lanceolate to lance-oblong), 3.2–5.5 mm, not enlarging in fruit, apex green to purple, acute, not hooded, glabrous;

petals oblanceolate to oblong-elliptic, 1–1.5 times as long as sepals, apex obtuse to rounded, entire.

Capsules

on stipe ca. 0.1 mm or shorter, narrowly ellipsoid, 3–4.2 mm, shorter than sepals.

on stipe about 0.2 mm, ± ovoid, 3.5–5.8 mm, equaling or longer than sepals.

Seeds

reddish brown to black, suborbiculate, radicle obscure, slightly compressed, 0.5–0.6 mm, tuberculate;

tubercles low, rounded.

brown or reddish, asymmetically reniform with radicle prolonged into beak, not compressed, 0.7–1 mm, minutely papillate.

Minuartia patula

Minuartia cismontana

Phenology Flowering spring–early summer. Flowering spring–summer.
Habitat Prairies, meadows, limestone barrens, and rocky outcrops in sandy, clayey, or gravelly soils Dry woodlands, chaparral, often on serpentine, (100-)400-1700 m
Elevation 0-500
Distribution
from FNA
AL; AR; GA; IL; IN; KS; KY; LA; MO; MS; OH; OK; PA; TN; TX; VA; WI
[BONAP county map]
from FNA
CA; OR
[BONAP county map]
Discussion

Minuartia patula and the related M. muscorum have received little attention in comparison to the granite-outcrop minuartias, the M. uniflora complex. J. A. Steyermark (1941) studied these taxa and described three forms, based chiefly on pubescence variation. Plants entirely glabrous [forma pitcheri (Nuttall) Steyermark] and those with sepals and pedicels somewhat stipitate-glandular (forma media Steyermark) were segregated from densely stipitate-glandular plants (forma patula). We do not feel that such variations deserve formal taxonomic recognition. Forma robusta, as defined by Steyermark, is here referred to M. muscorum.

Most specimens of Minuartia patula have prominently five-veined sepals (seen especially easily in the glabrous forms); occasional plants from Georgia, Kentucky, and Virginia have glabrous sepals with only three strong veins, resembling those of M. muscorum; in other features, including the seeds, they are clearly referable to M. patula. The status of the plants with three-veined sepals remains ambiguous; J. A. Steyermark (1941) included them in his forma media and B. Maguire (1951) included them (in our opinion incorrectly) in his var. robusta.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Minuartia cismontana is closely related to M. californica and M. pusilla, and has been overlooked as the former for many years. Phenology and elevation appear to segregate the species in areas of California where the ranges of M. cismontana and M. californica overlap (R. J. Meinke and P. F. Zika 1992).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 5, p. 131. FNA vol. 5, p. 122.
Parent taxa Caryophyllaceae > subfam. Alsinoideae > Minuartia Caryophyllaceae > subfam. Alsinoideae > Minuartia
Sibling taxa
M. arctica, M. austromontana, M. biflora, M. californica, M. caroliniana, M. cismontana, M. cumberlandensis, M. dawsonensis, M. decumbens, M. douglasii, M. drummondii, M. elegans, M. glabra, M. godfreyi, M. groenlandica, M. howellii, M. macrantha, M. macrocarpa, M. marcescens, M. michauxii, M. muscorum, M. nuttallii, M. obtusiloba, M. pusilla, M. rosei, M. rossii, M. rubella, M. stolonifera, M. stricta, M. tenella, M. uniflora, M. yukonensis
M. arctica, M. austromontana, M. biflora, M. californica, M. caroliniana, M. cumberlandensis, M. dawsonensis, M. decumbens, M. douglasii, M. drummondii, M. elegans, M. glabra, M. godfreyi, M. groenlandica, M. howellii, M. macrantha, M. macrocarpa, M. marcescens, M. michauxii, M. muscorum, M. nuttallii, M. obtusiloba, M. patula, M. pusilla, M. rosei, M. rossii, M. rubella, M. stolonifera, M. stricta, M. tenella, M. uniflora, M. yukonensis
Synonyms Arenaria patula, Alsinopsis patula, Alsinopsis pitcheri, Sabulina patula
Name authority (Michaux) Mattfeld: Bot. Jahrb. Syst. 57(Beibl. 126): 28. (1921) Meinke & Zika: Madroño 39: 289, figs. 1, 3. (1992)
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