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brittle sandwort, musk-flower, Nuttall sandwort, Nuttall's sandwort, Nuttall's stitchwort

Yukon stitchwort

Habit Plants perennial, mat-forming. Plants perennial, mat-forming.
Taproots

thickened, woody;

crown, many-branched, woody;

rhizomes and trailing stems to 60 cm.

stout, woody.

Stems

ascending to erect, ± green, 2–20 cm, densely glandular-hairy throughout, internodes of flowering stems 0.2–2 times as long as leaves.

erect to ascending, green, 10–30 cm, minutely retrorsely pubescent proximally, flowering stems stipitate-glandular, usually densely so, internodes of flowering stems 3–5 times as long as leaves.

Leaves

tightly appressed to spreading, ± evenly spaced, connate proximally, with ± loose, scarious sheath 0.1–0.7 mm;

blade straight to recurved, ± green, flat, prominently 1-veined abaxially, broadly lanceolate to linear, 5–20 × 0.5–1.5 mm, ± rigid, margins rounded, scarious in proximal 1/3–1/4, apex green to purple, acute to acuminate or spinescent, navicular with small mucro or spinescent, dull, stipitate-glandular;

axillary leaves present proximally to throughout.

tightly overlapping (vegetative), variably spaced (cauline), usually connate proximally, with tight, scarious to herbaceous sheath 0.5–1.8 mm;

blade straight to outwardly curved, green, flat, 3-veined abaxially, midvein more prominent than 2 lateral veins, filiform-linear, 10–18 × 0.8–1.3 mm, flexuous, margins not thickened, scarious, ciliate, often sparsely so, apex green, often with white callosity, acuminate-pungent, flat to slightly navicular, shiny, glabrous;

axillary leaves present among vegetative leaves.

Inflorescences

(3–)6–30-flowered, open cymes;

bracts lanceolate to subulate, usually scarious.

3–13-flowered, open cymes;

bracts lanceolate, herbaceous.

Pedicels

0.2–2 cm, stipitate-glandular.

2–5 cm, densely stipitate-glandular.

Flowers

hypanthium disc-shaped;

sepals 1–3-veined, narrowly lanceolate to lanceolate or ovate (herbaceous portion narrowly lanceolate to lanceolate or ovate), 3–6(–7) mm, not enlarging in fruit, apex often purple, acute to acuminate or spinescent, not hooded, stipitate-glandular;

petals obovate, 0.5–1.6 times as long as sepals, apex rounded, entire.

hypanthium cup-shaped;

sepals prominently 3-veined, lanceolate to oblong-lanceolate (herbaceous lanceolate to oblong-lanceolate), 6–8 mm, not enlarging in fruit, apex green to purplish, rounded, hooded, stipitate-glandular, especially proximally;

petals oblanceolate, 1.3–1.5 times as long as sepals, apex rounded, entire.

Capsules

on stipe ca. 0.1–0.2 mm, ovoid, 5 mm, usually shorter than sepals.

narrowly ellipsoid, 7–10 mm, longer than sepals.

Seeds

reddish brown to dark brown, oblong-elliptic with hilar notch on 1 end, 1.5–2.7 mm, tuberculate;

tubercles low-rounded.

reniform, ca. 1 mm, tuberculate.

Minuartia nuttallii

Minuartia yukonensis

Phenology Flowering spring–summer.
Habitat Dry, rocky slopes and meadows, scree slopes into alpine zone
Elevation 0-1000 m (0-3300 ft)
Distribution
from FNA
CA; CO; ID; MT; NV; OR; UT; WA; WY; AB; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; BC; NT; YT; Asia (Russian Far East)
[BONAP county map]
Discussion

Varieties 4 (4 in the flora).

Minuartia nuttallii, M. decumbens, M. rosei, and M. stolonifera form a complex that, together with the eastern species M. caroliniana and M. michauxii, comprise sect. Sclerophylla Mattfeld. The four western species all have capsules that contain one to three(?) large (1.5–2.8 mm) seeds; unfortunately, these plants appear to be collected only rarely in fruit.

Minuartia nuttallii includes four varieties, which can, for the most part, be easily recognized. There is some overlap between var. gracilis and var. fragilis in western Nevada and southeastern Oregon, where some plants exhibit prominently arcuate-spreading leaves (as in var. fragilis) and weakly veined sepals (as in var. gracilis).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Minuartia yukonensis is an amphi-Beringian species, reported from two sites in the Russian Far East. Some collections of it may be labeled as Arenaria laricifolia Linnaeus, a European species to which Alaskan material has been misattributed.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blade apices spinecent; sepal apices spinescent
→ 2
1. Leaf blade apices acute to acuminate, somewhat navicular; sepal apices acuminate
→ 3
2. Leaves prominently arcuate-spreading, blade 10-20 mm; sepals (1-)3-veined
var. fragilis
2. Leaves appressed to occasionally arcuate- spreading, blade 5-7 mm; sepals 1(-3)-veined
var. gracilis
3. Sepals lanceolate to narrowly so; petals 0.5-1.2 times as long as sepals
var. nuttallii
3. Sepals ovate to lanceolate; petals 1.1-1.6 times as long as sepals
var. gregaria
Source FNA vol. 5, p. 129. FNA vol. 5, p. 136.
Parent taxa Caryophyllaceae > subfam. Alsinoideae > Minuartia Caryophyllaceae > subfam. Alsinoideae > Minuartia
Sibling taxa
M. arctica, M. austromontana, M. biflora, M. californica, M. caroliniana, M. cismontana, M. cumberlandensis, M. dawsonensis, M. decumbens, M. douglasii, M. drummondii, M. elegans, M. glabra, M. godfreyi, M. groenlandica, M. howellii, M. macrantha, M. macrocarpa, M. marcescens, M. michauxii, M. muscorum, M. obtusiloba, M. patula, M. pusilla, M. rosei, M. rossii, M. rubella, M. stolonifera, M. stricta, M. tenella, M. uniflora, M. yukonensis
M. arctica, M. austromontana, M. biflora, M. californica, M. caroliniana, M. cismontana, M. cumberlandensis, M. dawsonensis, M. decumbens, M. douglasii, M. drummondii, M. elegans, M. glabra, M. godfreyi, M. groenlandica, M. howellii, M. macrantha, M. macrocarpa, M. marcescens, M. michauxii, M. muscorum, M. nuttallii, M. obtusiloba, M. patula, M. pusilla, M. rosei, M. rossii, M. rubella, M. stolonifera, M. stricta, M. tenella, M. uniflora
Subordinate taxa
M. nuttallii var. fragilis, M. nuttallii var. gracilis, M. nuttallii var. gregaria, M. nuttallii var. nuttallii
Synonyms Arenaria nuttallii, Minuopsis nuttallii Lidia yukonensis
Name authority (Pax) Briquet: Annuaire Conserv. Jard. Bot. Genève 13–14: 385. (1911) Hultén: Ark. Bot., n. s. 7: 521968
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