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brittle sandwort, musk-flower, Nuttall sandwort, Nuttall's sandwort, Nuttall's stitchwort

slender sandwort, slender stitchwort

Habit Plants perennial, mat-forming. Plants annual.
Taproots

thickened, woody;

crown, many-branched, woody;

rhizomes and trailing stems to 60 cm.

filiform.

Stems

ascending to erect, ± green, 2–20 cm, densely glandular-hairy throughout, internodes of flowering stems 0.2–2 times as long as leaves.

erect, green, 5–25 cm, stipitate-glandular distally or throughout, internodes of stems 2–5 times as long as leaves.

Leaves

tightly appressed to spreading, ± evenly spaced, connate proximally, with ± loose, scarious sheath 0.1–0.7 mm;

blade straight to recurved, ± green, flat, prominently 1-veined abaxially, broadly lanceolate to linear, 5–20 × 0.5–1.5 mm, ± rigid, margins rounded, scarious in proximal 1/3–1/4, apex green to purple, acute to acuminate or spinescent, navicular with small mucro or spinescent, dull, stipitate-glandular;

axillary leaves present proximally to throughout.

overlapping proximally, often connate basally, with loose, scarious sheath 0.2–0.5 mm;

blade straight to outwardly curved, green, flat to concave, prominently 1-veined abaxially, narrowly lanceolate to subulate, 5–17 × 0.5–1.5 mm, flexuous, margins not thickened, often scarious, sometimes ciliate or stipitate-glandular, apex purple, apiculate, navicular, shiny to dull, glabrous or stipitate-glandular;

axillary leaves often present.

Inflorescences

(3–)6–30-flowered, open cymes;

bracts lanceolate to subulate, usually scarious.

7–25+-flowered, open cymes;

bracts subulate to lanceolate, scarious.

Pedicels

0.2–2 cm, stipitate-glandular.

0.2–1.5 cm, stipitate-glandular.

Flowers

hypanthium disc-shaped;

sepals 1–3-veined, narrowly lanceolate to lanceolate or ovate (herbaceous portion narrowly lanceolate to lanceolate or ovate), 3–6(–7) mm, not enlarging in fruit, apex often purple, acute to acuminate or spinescent, not hooded, stipitate-glandular;

petals obovate, 0.5–1.6 times as long as sepals, apex rounded, entire.

hypanthium disc-shaped;

sepals prominently 3-veined, ovate to narrowly so (herbaceous portion narrowly ovate to lanceolate), 2.5–3 mm, not enlarging in fruit, apex green to purple, acute to acuminate, not hooded, densely stipitate-glandular;

petals obovate, 1.5–2 times as long as sepals, apex rounded, entire.

Capsules

on stipe ca. 0.1–0.2 mm, ovoid, 5 mm, usually shorter than sepals.

on stipe ca. 0.1 mm, ovoid, 3–4 mm, longer than sepals.

Seeds

reddish brown to dark brown, oblong-elliptic with hilar notch on 1 end, 1.5–2.7 mm, tuberculate;

tubercles low-rounded.

brown, suborbiculate with radicle prolonged to rounded beak, somewhat compressed, 0.4–0.6 mm, tuberculate;

tubercles low, rounded, elongate.

2n

= 24.

Minuartia nuttallii

Minuartia tenella

Phenology Flowering spring–summer.
Habitat Coastal bluffs and forest openings
Elevation 0-700 m (0-2300 ft)
Distribution
from FNA
CA; CO; ID; MT; NV; OR; UT; WA; WY; AB; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 4 (4 in the flora).

Minuartia nuttallii, M. decumbens, M. rosei, and M. stolonifera form a complex that, together with the eastern species M. caroliniana and M. michauxii, comprise sect. Sclerophylla Mattfeld. The four western species all have capsules that contain one to three(?) large (1.5–2.8 mm) seeds; unfortunately, these plants appear to be collected only rarely in fruit.

Minuartia nuttallii includes four varieties, which can, for the most part, be easily recognized. There is some overlap between var. gracilis and var. fragilis in western Nevada and southeastern Oregon, where some plants exhibit prominently arcuate-spreading leaves (as in var. fragilis) and weakly veined sepals (as in var. gracilis).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Although B. Maguire (1951, 1958) included Minuartia tenella within his concept of Arenaria stricta (M. michauxii), we see little more than a superficial resemblance between the taxa as we circumscribe them.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blade apices spinecent; sepal apices spinescent
→ 2
1. Leaf blade apices acute to acuminate, somewhat navicular; sepal apices acuminate
→ 3
2. Leaves prominently arcuate-spreading, blade 10-20 mm; sepals (1-)3-veined
var. fragilis
2. Leaves appressed to occasionally arcuate- spreading, blade 5-7 mm; sepals 1(-3)-veined
var. gracilis
3. Sepals lanceolate to narrowly so; petals 0.5-1.2 times as long as sepals
var. nuttallii
3. Sepals ovate to lanceolate; petals 1.1-1.6 times as long as sepals
var. gregaria
Source FNA vol. 5, p. 129. FNA vol. 5, p. 135.
Parent taxa Caryophyllaceae > subfam. Alsinoideae > Minuartia Caryophyllaceae > subfam. Alsinoideae > Minuartia
Sibling taxa
M. arctica, M. austromontana, M. biflora, M. californica, M. caroliniana, M. cismontana, M. cumberlandensis, M. dawsonensis, M. decumbens, M. douglasii, M. drummondii, M. elegans, M. glabra, M. godfreyi, M. groenlandica, M. howellii, M. macrantha, M. macrocarpa, M. marcescens, M. michauxii, M. muscorum, M. obtusiloba, M. patula, M. pusilla, M. rosei, M. rossii, M. rubella, M. stolonifera, M. stricta, M. tenella, M. uniflora, M. yukonensis
M. arctica, M. austromontana, M. biflora, M. californica, M. caroliniana, M. cismontana, M. cumberlandensis, M. dawsonensis, M. decumbens, M. douglasii, M. drummondii, M. elegans, M. glabra, M. godfreyi, M. groenlandica, M. howellii, M. macrantha, M. macrocarpa, M. marcescens, M. michauxii, M. muscorum, M. nuttallii, M. obtusiloba, M. patula, M. pusilla, M. rosei, M. rossii, M. rubella, M. stolonifera, M. stricta, M. uniflora, M. yukonensis
Subordinate taxa
M. nuttallii var. fragilis, M. nuttallii var. gracilis, M. nuttallii var. gregaria, M. nuttallii var. nuttallii
Synonyms Arenaria nuttallii, Minuopsis nuttallii Greniera tenella, Alsinopsis tenella, Arenaria macra, Arenaria stricta, Arenaria stricta subsp. macra, Arenaria stricta var. puberulenta
Name authority (Pax) Briquet: Annuaire Conserv. Jard. Bot. Genève 13–14: 385. (1911) (J. Gay) Mattfeld: Bot. Jahrb. Syst. 57(Beibl. 126): 29. (1921)
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