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brittle sandwort, musk-flower, Nuttall sandwort, Nuttall's sandwort, Nuttall's stitchwort

Michaux's sandplant, Michaux's stitchwort, rock sandwort

Habit Plants perennial, mat-forming. Plants perennial, cespitose, sometimes matted.
Taproots

thickened, woody;

crown, many-branched, woody;

rhizomes and trailing stems to 60 cm.

thickened, woody;

crown many-branched, thickened.

Stems

ascending to erect, ± green, 2–20 cm, densely glandular-hairy throughout, internodes of flowering stems 0.2–2 times as long as leaves.

erect to ascending, green, 8–40 cm, glabrous, internodes of flowering stems 0.5–10+ times as long as leaves.

Leaves

tightly appressed to spreading, ± evenly spaced, connate proximally, with ± loose, scarious sheath 0.1–0.7 mm;

blade straight to recurved, ± green, flat, prominently 1-veined abaxially, broadly lanceolate to linear, 5–20 × 0.5–1.5 mm, ± rigid, margins rounded, scarious in proximal 1/3–1/4, apex green to purple, acute to acuminate or spinescent, navicular with small mucro or spinescent, dull, stipitate-glandular;

axillary leaves present proximally to throughout.

tightly overlapping in proximal 1/3 of stem, variable spaced, usually connate proximally, with loose, scarious sheath 0.2–1 mm;

blade erect to spreading, green, flat or convex to 3-angled, 1–3-veined, prominently so abaxially, filiform, linear to linear-lanceolate, 8–30 × 0.5–1.8 mm, rigid, margins not thickened, ± scarious, smooth, apex green, blunt to pungent, flat to navicular, shiny, glabrous;

axillary leaves present among vegetative leaves.

Inflorescences

(3–)6–30-flowered, open cymes;

bracts lanceolate to subulate, usually scarious.

5–30-flowered, lax to congested cymes;

bracts narrowly lanceolate to subulate, herbaceous, margins scarious.

Pedicels

0.2–2 cm, stipitate-glandular.

0.3–6 cm, glabrous.

Flowers

hypanthium disc-shaped;

sepals 1–3-veined, narrowly lanceolate to lanceolate or ovate (herbaceous portion narrowly lanceolate to lanceolate or ovate), 3–6(–7) mm, not enlarging in fruit, apex often purple, acute to acuminate or spinescent, not hooded, stipitate-glandular;

petals obovate, 0.5–1.6 times as long as sepals, apex rounded, entire.

hypanthium dish-shaped;

sepals 3-veined, ovate to lanceolate (herbaceous portion ovate to narrowly lanceolate), 3–6 mm, not enlarging in fruit, apex green, acute to mostly acuminate, not hooded, glabrous;

petals oblong-obovate, 1.3–2 times as long as sepals, apex rounded, entire.

Capsules

on stipe ca. 0.1–0.2 mm, ovoid, 5 mm, usually shorter than sepals.

on stipe ca. 0.1–0.2 mm, ellipsoid, 3–4 mm, usually shorter than sepals.

Seeds

reddish brown to dark brown, oblong-elliptic with hilar notch on 1 end, 1.5–2.7 mm, tuberculate;

tubercles low-rounded.

black, suborbiculate, compressed, 0.8–0.9 mm, tuberculate;

tubercles elongate.

2n

= 30(?), 44.

Minuartia nuttallii

Minuartia michauxii

Phenology Flowering spring–summer.
Habitat Dry, calcareous gravel and ledges
Elevation 0-1000 m (0-3300 ft)
Distribution
from FNA
CA; CO; ID; MT; NV; OR; UT; WA; WY; AB; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; DE; IA; IL; IN; KS; KY; MA; MD; MI; MO; NE; NH; NJ; NM; NY; OH; OK; PA; RI; SD; TX; VA; VT; WI; WV; ON; QC
[BONAP county map]
Discussion

Varieties 4 (4 in the flora).

Minuartia nuttallii, M. decumbens, M. rosei, and M. stolonifera form a complex that, together with the eastern species M. caroliniana and M. michauxii, comprise sect. Sclerophylla Mattfeld. The four western species all have capsules that contain one to three(?) large (1.5–2.8 mm) seeds; unfortunately, these plants appear to be collected only rarely in fruit.

Minuartia nuttallii includes four varieties, which can, for the most part, be easily recognized. There is some overlap between var. gracilis and var. fragilis in western Nevada and southeastern Oregon, where some plants exhibit prominently arcuate-spreading leaves (as in var. fragilis) and weakly veined sepals (as in var. gracilis).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

We concur with J. A. Steyermark (1963) and G. Yatskievych and J. Turner (1990) that the concept of Arenaria texana originally put forth by Britton requires further study. Plants labeled as Minuartia michauxii var. texana have slightly shorter leaves that are crowded into the proximal one-third rather than one-half of the somewhat shorter stem. The plants are often in the southern part of the range and may, as Steyermark noted, be associated with more open, xeric habitats.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blade apices spinecent; sepal apices spinescent
→ 2
1. Leaf blade apices acute to acuminate, somewhat navicular; sepal apices acuminate
→ 3
2. Leaves prominently arcuate-spreading, blade 10-20 mm; sepals (1-)3-veined
var. fragilis
2. Leaves appressed to occasionally arcuate- spreading, blade 5-7 mm; sepals 1(-3)-veined
var. gracilis
3. Sepals lanceolate to narrowly so; petals 0.5-1.2 times as long as sepals
var. nuttallii
3. Sepals ovate to lanceolate; petals 1.1-1.6 times as long as sepals
var. gregaria
Source FNA vol. 5, p. 129. FNA vol. 5, p. 128.
Parent taxa Caryophyllaceae > subfam. Alsinoideae > Minuartia Caryophyllaceae > subfam. Alsinoideae > Minuartia
Sibling taxa
M. arctica, M. austromontana, M. biflora, M. californica, M. caroliniana, M. cismontana, M. cumberlandensis, M. dawsonensis, M. decumbens, M. douglasii, M. drummondii, M. elegans, M. glabra, M. godfreyi, M. groenlandica, M. howellii, M. macrantha, M. macrocarpa, M. marcescens, M. michauxii, M. muscorum, M. obtusiloba, M. patula, M. pusilla, M. rosei, M. rossii, M. rubella, M. stolonifera, M. stricta, M. tenella, M. uniflora, M. yukonensis
M. arctica, M. austromontana, M. biflora, M. californica, M. caroliniana, M. cismontana, M. cumberlandensis, M. dawsonensis, M. decumbens, M. douglasii, M. drummondii, M. elegans, M. glabra, M. godfreyi, M. groenlandica, M. howellii, M. macrantha, M. macrocarpa, M. marcescens, M. muscorum, M. nuttallii, M. obtusiloba, M. patula, M. pusilla, M. rosei, M. rossii, M. rubella, M. stolonifera, M. stricta, M. tenella, M. uniflora, M. yukonensis
Subordinate taxa
M. nuttallii var. fragilis, M. nuttallii var. gracilis, M. nuttallii var. gregaria, M. nuttallii var. nuttallii
Synonyms Arenaria nuttallii, Minuopsis nuttallii Alsine michauxii, Arenaria stricta subsp. texana, Arenaria stricta var. texana, Arenaria texana, M. michauxii var. texana, Minuopsis michauxii, Sabulina stricta
Name authority (Pax) Briquet: Annuaire Conserv. Jard. Bot. Genève 13–14: 385. (1911) (Fenzl) Farwell: Rep. (Annual) Michigan Acad. Sci. 20: 177. (1919)
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