Minuartia nuttallii |
Minuartia michauxii |
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brittle sandwort, musk-flower, Nuttall sandwort, Nuttall's sandwort, Nuttall's stitchwort |
Michaux's sandplant, Michaux's stitchwort, rock sandwort |
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Habit | Plants perennial, mat-forming. | Plants perennial, cespitose, sometimes matted. | ||||||||||||
Taproots | thickened, woody; crown, many-branched, woody; rhizomes and trailing stems to 60 cm. |
thickened, woody; crown many-branched, thickened. |
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Stems | ascending to erect, ± green, 2–20 cm, densely glandular-hairy throughout, internodes of flowering stems 0.2–2 times as long as leaves. |
erect to ascending, green, 8–40 cm, glabrous, internodes of flowering stems 0.5–10+ times as long as leaves. |
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Leaves | tightly appressed to spreading, ± evenly spaced, connate proximally, with ± loose, scarious sheath 0.1–0.7 mm; blade straight to recurved, ± green, flat, prominently 1-veined abaxially, broadly lanceolate to linear, 5–20 × 0.5–1.5 mm, ± rigid, margins rounded, scarious in proximal 1/3–1/4, apex green to purple, acute to acuminate or spinescent, navicular with small mucro or spinescent, dull, stipitate-glandular; axillary leaves present proximally to throughout. |
tightly overlapping in proximal 1/3 of stem, variable spaced, usually connate proximally, with loose, scarious sheath 0.2–1 mm; blade erect to spreading, green, flat or convex to 3-angled, 1–3-veined, prominently so abaxially, filiform, linear to linear-lanceolate, 8–30 × 0.5–1.8 mm, rigid, margins not thickened, ± scarious, smooth, apex green, blunt to pungent, flat to navicular, shiny, glabrous; axillary leaves present among vegetative leaves. |
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Inflorescences | (3–)6–30-flowered, open cymes; bracts lanceolate to subulate, usually scarious. |
5–30-flowered, lax to congested cymes; bracts narrowly lanceolate to subulate, herbaceous, margins scarious. |
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Pedicels | 0.2–2 cm, stipitate-glandular. |
0.3–6 cm, glabrous. |
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Flowers | hypanthium disc-shaped; sepals 1–3-veined, narrowly lanceolate to lanceolate or ovate (herbaceous portion narrowly lanceolate to lanceolate or ovate), 3–6(–7) mm, not enlarging in fruit, apex often purple, acute to acuminate or spinescent, not hooded, stipitate-glandular; petals obovate, 0.5–1.6 times as long as sepals, apex rounded, entire. |
hypanthium dish-shaped; sepals 3-veined, ovate to lanceolate (herbaceous portion ovate to narrowly lanceolate), 3–6 mm, not enlarging in fruit, apex green, acute to mostly acuminate, not hooded, glabrous; petals oblong-obovate, 1.3–2 times as long as sepals, apex rounded, entire. |
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Capsules | on stipe ca. 0.1–0.2 mm, ovoid, 5 mm, usually shorter than sepals. |
on stipe ca. 0.1–0.2 mm, ellipsoid, 3–4 mm, usually shorter than sepals. |
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Seeds | reddish brown to dark brown, oblong-elliptic with hilar notch on 1 end, 1.5–2.7 mm, tuberculate; tubercles low-rounded. |
black, suborbiculate, compressed, 0.8–0.9 mm, tuberculate; tubercles elongate. |
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2n | = 30(?), 44. |
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Minuartia nuttallii |
Minuartia michauxii |
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Phenology | Flowering spring–summer. | |||||||||||||
Habitat | Dry, calcareous gravel and ledges | |||||||||||||
Elevation | 0-1000 m (0-3300 ft) | |||||||||||||
Distribution |
CA; CO; ID; MT; NV; OR; UT; WA; WY; AB; BC
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AL; AR; CT; DE; IA; IL; IN; KS; KY; MA; MD; MI; MO; NE; NH; NJ; NM; NY; OH; OK; PA; RI; SD; TX; VA; VT; WI; WV; ON; QC |
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Discussion | Varieties 4 (4 in the flora). Minuartia nuttallii, M. decumbens, M. rosei, and M. stolonifera form a complex that, together with the eastern species M. caroliniana and M. michauxii, comprise sect. Sclerophylla Mattfeld. The four western species all have capsules that contain one to three(?) large (1.5–2.8 mm) seeds; unfortunately, these plants appear to be collected only rarely in fruit. Minuartia nuttallii includes four varieties, which can, for the most part, be easily recognized. There is some overlap between var. gracilis and var. fragilis in western Nevada and southeastern Oregon, where some plants exhibit prominently arcuate-spreading leaves (as in var. fragilis) and weakly veined sepals (as in var. gracilis). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
We concur with J. A. Steyermark (1963) and G. Yatskievych and J. Turner (1990) that the concept of Arenaria texana originally put forth by Britton requires further study. Plants labeled as Minuartia michauxii var. texana have slightly shorter leaves that are crowded into the proximal one-third rather than one-half of the somewhat shorter stem. The plants are often in the southern part of the range and may, as Steyermark noted, be associated with more open, xeric habitats. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 5, p. 129. | FNA vol. 5, p. 128. | ||||||||||||
Parent taxa | Caryophyllaceae > subfam. Alsinoideae > Minuartia | Caryophyllaceae > subfam. Alsinoideae > Minuartia | ||||||||||||
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Subordinate taxa | ||||||||||||||
Synonyms | Arenaria nuttallii, Minuopsis nuttallii | Alsine michauxii, Arenaria stricta subsp. texana, Arenaria stricta var. texana, Arenaria texana, M. michauxii var. texana, Minuopsis michauxii, Sabulina stricta | ||||||||||||
Name authority | (Pax) Briquet: Annuaire Conserv. Jard. Bot. Genève 13–14: 385. (1911) | (Fenzl) Farwell: Rep. (Annual) Michigan Acad. Sci. 20: 177. (1919) | ||||||||||||
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