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brittle sandwort, musk-flower, Nuttall sandwort, Nuttall's sandwort, Nuttall's stitchwort

minuartie de Dawson, rock stitchwort

Habit Plants perennial, mat-forming. Plants perennial, sometimes mat-forming, green.
Taproots

thickened, woody;

crown, many-branched, woody;

rhizomes and trailing stems to 60 cm.

filiform to somewhat thickened.

Stems

ascending to erect, ± green, 2–20 cm, densely glandular-hairy throughout, internodes of flowering stems 0.2–2 times as long as leaves.

erect to ascending, green, 4–30 cm, glabrous, internodes of flowering stems 1–10 times as long as leaves.

Leaves

tightly appressed to spreading, ± evenly spaced, connate proximally, with ± loose, scarious sheath 0.1–0.7 mm;

blade straight to recurved, ± green, flat, prominently 1-veined abaxially, broadly lanceolate to linear, 5–20 × 0.5–1.5 mm, ± rigid, margins rounded, scarious in proximal 1/3–1/4, apex green to purple, acute to acuminate or spinescent, navicular with small mucro or spinescent, dull, stipitate-glandular;

axillary leaves present proximally to throughout.

overlapping or crowded proximally, variably spaced distally, connate proximally, with tight, scarious to herbaceous sheath 0.2–0.5 mm;

blade straight to slightly outwardly curved, green, flat, 1-veined, occasionally 3-veined abaxially, linear to subulate, 4–15 × 0.5–2 mm, flexuous, margins not thickened, scarious, smooth, apex green or purple, mostly rounded, slightly navicular, shiny, glabrous;

axillary leaves present among proximal cauline leaves.

Inflorescences

(3–)6–30-flowered, open cymes;

bracts lanceolate to subulate, usually scarious.

7–15-flowered (rarely fewer), open cymes;

bracts subulate, herbaceous.

Pedicels

0.2–2 cm, stipitate-glandular.

0.3–2.5 cm, glabrous.

Flowers

hypanthium disc-shaped;

sepals 1–3-veined, narrowly lanceolate to lanceolate or ovate (herbaceous portion narrowly lanceolate to lanceolate or ovate), 3–6(–7) mm, not enlarging in fruit, apex often purple, acute to acuminate or spinescent, not hooded, stipitate-glandular;

petals obovate, 0.5–1.6 times as long as sepals, apex rounded, entire.

hypanthium disc-shaped;

sepals prominently 3-veined, ovate to broadly lanceolate (herbaceous portion ovate to broadly lanceolate), 2.5–3.2 mm, to 4 mm in fruit, apex green to purple, acute to apiculate, not hooded, glabrous;

petals lancolate to spatulate, 0.5–0.8 times as long as sepals, apex rounded, entire, or petals absent.

Capsules

on stipe ca. 0.1–0.2 mm, ovoid, 5 mm, usually shorter than sepals.

on stipe ca. 0.2 mm, ovoid, 3.5–4.5 mm, longer than sepals.

Seeds

reddish brown to dark brown, oblong-elliptic with hilar notch on 1 end, 1.5–2.7 mm, tuberculate;

tubercles low-rounded.

dark brown to black, suborbiculate with radicle prolonged into tiny beak, 0.5–0.6 mm, tuberculate;

tubercles low, rounded, somewhat elongate.

2n

= 30, 60.

Minuartia nuttallii

Minuartia dawsonensis

Phenology Flowering late spring–summer.
Habitat Moist, calcareous ledges and gravelly areas (dry, open, and sometimes disturbed slopes, calcareous-gravel raised beach ridges, thin soil over limestone) in mesic forest openings and meadows in montane and subalpine areas and boreal plains, dry, open outcrops in oak or juniper savannas or prairies
Elevation 0-900 m (0-3000 ft)
Distribution
from FNA
CA; CO; ID; MT; NV; OR; UT; WA; WY; AB; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; MI; MN; ND; WI; AB; BC; MB; NL; NT; NU; ON; QC; SK; YT
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 4 (4 in the flora).

Minuartia nuttallii, M. decumbens, M. rosei, and M. stolonifera form a complex that, together with the eastern species M. caroliniana and M. michauxii, comprise sect. Sclerophylla Mattfeld. The four western species all have capsules that contain one to three(?) large (1.5–2.8 mm) seeds; unfortunately, these plants appear to be collected only rarely in fruit.

Minuartia nuttallii includes four varieties, which can, for the most part, be easily recognized. There is some overlap between var. gracilis and var. fragilis in western Nevada and southeastern Oregon, where some plants exhibit prominently arcuate-spreading leaves (as in var. fragilis) and weakly veined sepals (as in var. gracilis).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Although sometimes included in Minuartia michauxii [e.g., H. J. Scoggan’s (1978–1979, part 3) treatment of Arenaria stricta], M. dawsonensis is more closely related to the circumpolar M. stricta.

Minuartia litorea, known from Quebec and Ontario, may deserve recognition. An unpublished chromosome count suggests that it is a recent allopolyploid derived from M. dawsonensis and M. rubella (L. Brouillet, pers. comm.).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blade apices spinecent; sepal apices spinescent
→ 2
1. Leaf blade apices acute to acuminate, somewhat navicular; sepal apices acuminate
→ 3
2. Leaves prominently arcuate-spreading, blade 10-20 mm; sepals (1-)3-veined
var. fragilis
2. Leaves appressed to occasionally arcuate- spreading, blade 5-7 mm; sepals 1(-3)-veined
var. gracilis
3. Sepals lanceolate to narrowly so; petals 0.5-1.2 times as long as sepals
var. nuttallii
3. Sepals ovate to lanceolate; petals 1.1-1.6 times as long as sepals
var. gregaria
Source FNA vol. 5, p. 129. FNA vol. 5, p. 123.
Parent taxa Caryophyllaceae > subfam. Alsinoideae > Minuartia Caryophyllaceae > subfam. Alsinoideae > Minuartia
Sibling taxa
M. arctica, M. austromontana, M. biflora, M. californica, M. caroliniana, M. cismontana, M. cumberlandensis, M. dawsonensis, M. decumbens, M. douglasii, M. drummondii, M. elegans, M. glabra, M. godfreyi, M. groenlandica, M. howellii, M. macrantha, M. macrocarpa, M. marcescens, M. michauxii, M. muscorum, M. obtusiloba, M. patula, M. pusilla, M. rosei, M. rossii, M. rubella, M. stolonifera, M. stricta, M. tenella, M. uniflora, M. yukonensis
M. arctica, M. austromontana, M. biflora, M. californica, M. caroliniana, M. cismontana, M. cumberlandensis, M. decumbens, M. douglasii, M. drummondii, M. elegans, M. glabra, M. godfreyi, M. groenlandica, M. howellii, M. macrantha, M. macrocarpa, M. marcescens, M. michauxii, M. muscorum, M. nuttallii, M. obtusiloba, M. patula, M. pusilla, M. rosei, M. rossii, M. rubella, M. stolonifera, M. stricta, M. tenella, M. uniflora, M. yukonensis
Subordinate taxa
M. nuttallii var. fragilis, M. nuttallii var. gracilis, M. nuttallii var. gregaria, M. nuttallii var. nuttallii
Synonyms Arenaria nuttallii, Minuopsis nuttallii Arenaria dawsonensis, Alsinopsis dawsonensis, Arenaria litorea, Arenaria stricta var. dawsonensis, Arenaria stricta var. litorea, M. litorea, Sabulina dawsonensis
Name authority (Pax) Briquet: Annuaire Conserv. Jard. Bot. Genève 13–14: 385. (1911) (Britton) House: Amer. Midl. Naturalist 7: 132. (1921)
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