Minuartia nuttallii |
Minuartia cismontana |
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brittle sandwort, musk-flower, Nuttall sandwort, Nuttall's sandwort, Nuttall's stitchwort |
cismontane minuartia |
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Habit | Plants perennial, mat-forming. | Plants annual. | ||||||||||||
Taproots | thickened, woody; crown, many-branched, woody; rhizomes and trailing stems to 60 cm. |
filiform. |
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Stems | ascending to erect, ± green, 2–20 cm, densely glandular-hairy throughout, internodes of flowering stems 0.2–2 times as long as leaves. |
erect, green or reddish purple, (5–)8–20(–25) cm, glabrous, internodes of all stems 5–7 times as long as leaves. |
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Leaves | tightly appressed to spreading, ± evenly spaced, connate proximally, with ± loose, scarious sheath 0.1–0.7 mm; blade straight to recurved, ± green, flat, prominently 1-veined abaxially, broadly lanceolate to linear, 5–20 × 0.5–1.5 mm, ± rigid, margins rounded, scarious in proximal 1/3–1/4, apex green to purple, acute to acuminate or spinescent, navicular with small mucro or spinescent, dull, stipitate-glandular; axillary leaves present proximally to throughout. |
not overlapping, connate proximally, with loose, scarious sheath 0.1–0.3 mm; blade green or reddish purple, 3-veined proximally, midvein prominent abaxially, lateral veins 0.2–0.25 times as long as blade, straight to outwardly curved, flat, lance-attenuate to linear, 2–7(–9) × 0.5–1.2(–1.8) mm, flexuous, margins not thickened, scarious proximally, smooth, apex green or purple, rounded to acute, often mucronate, shiny, glabrous; axillary leaves occasionally present. |
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Inflorescences | (3–)6–30-flowered, open cymes; bracts lanceolate to subulate, usually scarious. |
5–20-flowered, open cymes; bracts subulate, herbaceous, often scarious-margined proximally. |
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Pedicels | 0.2–2 cm, stipitate-glandular. |
(0.7–)1–3(–3.5) cm, glabrous. |
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Flowers | hypanthium disc-shaped; sepals 1–3-veined, narrowly lanceolate to lanceolate or ovate (herbaceous portion narrowly lanceolate to lanceolate or ovate), 3–6(–7) mm, not enlarging in fruit, apex often purple, acute to acuminate or spinescent, not hooded, stipitate-glandular; petals obovate, 0.5–1.6 times as long as sepals, apex rounded, entire. |
hypanthium disc-shaped; sepals strongly 3(–5)-veined, lance-linear to lanceolate (herbaceous portion narrowly lanceolate to lance-oblong), 3.2–5.5 mm, not enlarging in fruit, apex green to purple, acute, not hooded, glabrous; petals oblanceolate to oblong-elliptic, 1–1.5 times as long as sepals, apex obtuse to rounded, entire. |
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Capsules | on stipe ca. 0.1–0.2 mm, ovoid, 5 mm, usually shorter than sepals. |
on stipe about 0.2 mm, ± ovoid, 3.5–5.8 mm, equaling or longer than sepals. |
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Seeds | reddish brown to dark brown, oblong-elliptic with hilar notch on 1 end, 1.5–2.7 mm, tuberculate; tubercles low-rounded. |
brown or reddish, asymmetically reniform with radicle prolonged into beak, not compressed, 0.7–1 mm, minutely papillate. |
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Minuartia nuttallii |
Minuartia cismontana |
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Phenology | Flowering spring–summer. | |||||||||||||
Habitat | Dry woodlands, chaparral, often on serpentine, (100-)400-1700 m | |||||||||||||
Distribution |
CA; CO; ID; MT; NV; OR; UT; WA; WY; AB; BC
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CA; OR |
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Discussion | Varieties 4 (4 in the flora). Minuartia nuttallii, M. decumbens, M. rosei, and M. stolonifera form a complex that, together with the eastern species M. caroliniana and M. michauxii, comprise sect. Sclerophylla Mattfeld. The four western species all have capsules that contain one to three(?) large (1.5–2.8 mm) seeds; unfortunately, these plants appear to be collected only rarely in fruit. Minuartia nuttallii includes four varieties, which can, for the most part, be easily recognized. There is some overlap between var. gracilis and var. fragilis in western Nevada and southeastern Oregon, where some plants exhibit prominently arcuate-spreading leaves (as in var. fragilis) and weakly veined sepals (as in var. gracilis). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Minuartia cismontana is closely related to M. californica and M. pusilla, and has been overlooked as the former for many years. Phenology and elevation appear to segregate the species in areas of California where the ranges of M. cismontana and M. californica overlap (R. J. Meinke and P. F. Zika 1992). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 5, p. 129. | FNA vol. 5, p. 122. | ||||||||||||
Parent taxa | Caryophyllaceae > subfam. Alsinoideae > Minuartia | Caryophyllaceae > subfam. Alsinoideae > Minuartia | ||||||||||||
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Subordinate taxa | ||||||||||||||
Synonyms | Arenaria nuttallii, Minuopsis nuttallii | |||||||||||||
Name authority | (Pax) Briquet: Annuaire Conserv. Jard. Bot. Genève 13–14: 385. (1911) | Meinke & Zika: Madroño 39: 289, figs. 1, 3. (1992) | ||||||||||||
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