Minuartia nuttallii |
Minuartia biflora |
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brittle sandwort, musk-flower, Nuttall sandwort, Nuttall's sandwort, Nuttall's stitchwort |
minuartie à deux fleurs, mountain stitchwort, mountain stitchwort or sandwort |
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Habit | Plants perennial, mat-forming. | Plants perennial, mat-forming. | ||||||||||||
Taproots | thickened, woody; crown, many-branched, woody; rhizomes and trailing stems to 60 cm. |
stout, woody. |
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Stems | ascending to erect, ± green, 2–20 cm, densely glandular-hairy throughout, internodes of flowering stems 0.2–2 times as long as leaves. |
suberect to ascending, green, 2–10-cm, retrorsely pubescent in lines, internodes of flowering stems 2–7 times as long as leaves. |
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Leaves | tightly appressed to spreading, ± evenly spaced, connate proximally, with ± loose, scarious sheath 0.1–0.7 mm; blade straight to recurved, ± green, flat, prominently 1-veined abaxially, broadly lanceolate to linear, 5–20 × 0.5–1.5 mm, ± rigid, margins rounded, scarious in proximal 1/3–1/4, apex green to purple, acute to acuminate or spinescent, navicular with small mucro or spinescent, dull, stipitate-glandular; axillary leaves present proximally to throughout. |
tightly overlapping or not (vegetative and proximal cauline) or variably spaced (distal cauline), usually connate proximally, with tight, scarious to herbaceous sheath 0.5–1 mm; blade straight to outwardly curved, green, flat, obscurely 1-veined abaxially, oblong or spatulate to elliptic, 5–10 × 0.7–2 mm, flexuous, margins not thickened, scarious, rarely ciliate proximally, apex green or purple, rounded, flat to navicular, shiny, glabrous; axillary leaves mostly absent. |
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Inflorescences | (3–)6–30-flowered, open cymes; bracts lanceolate to subulate, usually scarious. |
3–5-flowered, open cymes; bracts lanceolate, herbaceous. |
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Pedicels | 0.2–2 cm, stipitate-glandular. |
0.5–1 cm, usually densely stipitate-glandular. |
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Flowers | hypanthium disc-shaped; sepals 1–3-veined, narrowly lanceolate to lanceolate or ovate (herbaceous portion narrowly lanceolate to lanceolate or ovate), 3–6(–7) mm, not enlarging in fruit, apex often purple, acute to acuminate or spinescent, not hooded, stipitate-glandular; petals obovate, 0.5–1.6 times as long as sepals, apex rounded, entire. |
hypanthium cup-shaped; sepals 3-veined prominently in fruit, oblong to narrowly lanceolate (herbaceous portion often purple, oblong to narrowly oblanceolate), 3.5–4.5 mm, not enlarging in fruit, apex rarely purple, rounded, hooded (at least inner sepals) or not, glabrous to stipitate-glandular proximally; petals white or often lilac, broadly oblanceolate, 1.4–1.7 times as long as sepals, apex truncate, often shallowly notched. |
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Capsules | on stipe ca. 0.1–0.2 mm, ovoid, 5 mm, usually shorter than sepals. |
broadly ellipsoid, 5.5 mm, longer than sepals. |
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Seeds | reddish brown to dark brown, oblong-elliptic with hilar notch on 1 end, 1.5–2.7 mm, tuberculate; tubercles low-rounded. |
brown, suborbiculate with radicle prolonged into beak, slightly compressed, 0.7–0.8 mm, smooth or obscurely scupltured (50x). |
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2n | = 26. |
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Minuartia nuttallii |
Minuartia biflora |
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Phenology | Flowering spring–summer. | |||||||||||||
Habitat | Dry, calcareous, gravelly to rocky slopes, fell-fields, snow beds, heath in low arctic to alpine areas | |||||||||||||
Elevation | 0-2500 m (0-8200 ft) | |||||||||||||
Distribution |
CA; CO; ID; MT; NV; OR; UT; WA; WY; AB; BC
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AK; AB; BC; LB; NT; NU; QC; YT; Greenland; circumpolar; Europe; Asia
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Discussion | Varieties 4 (4 in the flora). Minuartia nuttallii, M. decumbens, M. rosei, and M. stolonifera form a complex that, together with the eastern species M. caroliniana and M. michauxii, comprise sect. Sclerophylla Mattfeld. The four western species all have capsules that contain one to three(?) large (1.5–2.8 mm) seeds; unfortunately, these plants appear to be collected only rarely in fruit. Minuartia nuttallii includes four varieties, which can, for the most part, be easily recognized. There is some overlap between var. gracilis and var. fragilis in western Nevada and southeastern Oregon, where some plants exhibit prominently arcuate-spreading leaves (as in var. fragilis) and weakly veined sepals (as in var. gracilis). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Specimens labeled Arenaria sajanensis Willdenow ex Schlechtendal from western North America, although sometimes referred to M. biflora (e.g., H. J. Scoggan 1978–1979, part 3), are likely to be M. obtusiloba. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 5, p. 129. | FNA vol. 5, p. 121. | ||||||||||||
Parent taxa | Caryophyllaceae > subfam. Alsinoideae > Minuartia | Caryophyllaceae > subfam. Alsinoideae > Minuartia | ||||||||||||
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Synonyms | Arenaria nuttallii, Minuopsis nuttallii | Stellaria biflora, Alsinopsis biflora, Lidia biflora | ||||||||||||
Name authority | (Pax) Briquet: Annuaire Conserv. Jard. Bot. Genève 13–14: 385. (1911) | (Linnaeus) Schinz & Thellung: Bull. Herb. Boissier, sér. 2, 7: 404. (1907) | ||||||||||||
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