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brittle sandwort, musk-flower, Nuttall sandwort, Nuttall's sandwort, Nuttall's stitchwort

arctic stitchwort, arctic stitchwort or sandwort

Habit Plants perennial, mat-forming. Plants perennial, mat-forming.
Taproots

thickened, woody;

crown, many-branched, woody;

rhizomes and trailing stems to 60 cm.

stout, woody.

Stems

ascending to erect, ± green, 2–20 cm, densely glandular-hairy throughout, internodes of flowering stems 0.2–2 times as long as leaves.

erect to ascending, green, 3–10 cm, retrorsely puberulent or stipitate-glandular, internodes of flowering stems 2–6 times as long as leaves.

Leaves

tightly appressed to spreading, ± evenly spaced, connate proximally, with ± loose, scarious sheath 0.1–0.7 mm;

blade straight to recurved, ± green, flat, prominently 1-veined abaxially, broadly lanceolate to linear, 5–20 × 0.5–1.5 mm, ± rigid, margins rounded, scarious in proximal 1/3–1/4, apex green to purple, acute to acuminate or spinescent, navicular with small mucro or spinescent, dull, stipitate-glandular;

axillary leaves present proximally to throughout.

tightly overlapping (vegetative) or variably spaced (cauline), usually connate proximally, with tight, scarious to herbaceous sheath 1.2–1.5 mm;

blade straight to outwardly curved, green, obscurely 1-veined, linear (proximal vegetative) or subulate (cauline), rounded 3-angled (abaxial surface thickened, rounded, adaxial surface flat to concave), 5–20 × 0.4–1 mm, flexuous, margins not thickened, herbaceous, often ciliate, apex often purple, rounded to truncate, shiny, glabrous (vegetative) or glabrous to stipitate-glandular (cauline);

axillary leaves absent.

Inflorescences

(3–)6–30-flowered, open cymes;

bracts lanceolate to subulate, usually scarious.

flowers solitary, terminal;

bracts narrowly lanceolate to oblong, herbaceous.

Pedicels

0.2–2 cm, stipitate-glandular.

0.5–3 cm, usually densely stipitate-glandular.

Flowers

hypanthium disc-shaped;

sepals 1–3-veined, narrowly lanceolate to lanceolate or ovate (herbaceous portion narrowly lanceolate to lanceolate or ovate), 3–6(–7) mm, not enlarging in fruit, apex often purple, acute to acuminate or spinescent, not hooded, stipitate-glandular;

petals obovate, 0.5–1.6 times as long as sepals, apex rounded, entire.

hypanthium cup-shaped;

sepals prominently 3-veined proximally, lanceolate to narrowly ovate (herbaceous portion often purple, ovate to oblong), 4–8 mm, enlarging slightly in fruit, apex often purple, rounded, hooded, stipitate-glandular;

petals oblanceolate, 1.5–2 times as long as sepals, apex broadly rounded, entire.

Capsules

on stipe ca. 0.1–0.2 mm, ovoid, 5 mm, usually shorter than sepals.

narrowly ellipsoid, 9–10 mm, longer than sepals.

Seeds

reddish brown to dark brown, oblong-elliptic with hilar notch on 1 end, 1.5–2.7 mm, tuberculate;

tubercles low-rounded.

brown, suborbiculate with radicle prolonged into beak, compressed, 1.2–1.6 mm, minutely tuberculate (50x).

2n

= 22 (Russia), 26 (Russia), 38 (Russia), 52, ca. 80.

Minuartia nuttallii

Minuartia arctica

Phenology Flowering summer.
Habitat Dry ridges, rocky mountain slopes, heathlands, alpine snowbed slopes, stony tundra
Elevation 0-1000 m (0-3300 ft)
Distribution
from FNA
CA; CO; ID; MT; NV; OR; UT; WA; WY; AB; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; NT; YT; Asia
[BONAP county map]
Discussion

Varieties 4 (4 in the flora).

Minuartia nuttallii, M. decumbens, M. rosei, and M. stolonifera form a complex that, together with the eastern species M. caroliniana and M. michauxii, comprise sect. Sclerophylla Mattfeld. The four western species all have capsules that contain one to three(?) large (1.5–2.8 mm) seeds; unfortunately, these plants appear to be collected only rarely in fruit.

Minuartia nuttallii includes four varieties, which can, for the most part, be easily recognized. There is some overlap between var. gracilis and var. fragilis in western Nevada and southeastern Oregon, where some plants exhibit prominently arcuate-spreading leaves (as in var. fragilis) and weakly veined sepals (as in var. gracilis).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Minuartia arctica is an amphi-Beringian species that is known to intergrade with M. obtusiloba. Hybrids between M. arctica and M. macrocarpa are known as well.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blade apices spinecent; sepal apices spinescent
→ 2
1. Leaf blade apices acute to acuminate, somewhat navicular; sepal apices acuminate
→ 3
2. Leaves prominently arcuate-spreading, blade 10-20 mm; sepals (1-)3-veined
var. fragilis
2. Leaves appressed to occasionally arcuate- spreading, blade 5-7 mm; sepals 1(-3)-veined
var. gracilis
3. Sepals lanceolate to narrowly so; petals 0.5-1.2 times as long as sepals
var. nuttallii
3. Sepals ovate to lanceolate; petals 1.1-1.6 times as long as sepals
var. gregaria
Source FNA vol. 5, p. 129. FNA vol. 5, p. 120.
Parent taxa Caryophyllaceae > subfam. Alsinoideae > Minuartia Caryophyllaceae > subfam. Alsinoideae > Minuartia
Sibling taxa
M. arctica, M. austromontana, M. biflora, M. californica, M. caroliniana, M. cismontana, M. cumberlandensis, M. dawsonensis, M. decumbens, M. douglasii, M. drummondii, M. elegans, M. glabra, M. godfreyi, M. groenlandica, M. howellii, M. macrantha, M. macrocarpa, M. marcescens, M. michauxii, M. muscorum, M. obtusiloba, M. patula, M. pusilla, M. rosei, M. rossii, M. rubella, M. stolonifera, M. stricta, M. tenella, M. uniflora, M. yukonensis
M. austromontana, M. biflora, M. californica, M. caroliniana, M. cismontana, M. cumberlandensis, M. dawsonensis, M. decumbens, M. douglasii, M. drummondii, M. elegans, M. glabra, M. godfreyi, M. groenlandica, M. howellii, M. macrantha, M. macrocarpa, M. marcescens, M. michauxii, M. muscorum, M. nuttallii, M. obtusiloba, M. patula, M. pusilla, M. rosei, M. rossii, M. rubella, M. stolonifera, M. stricta, M. tenella, M. uniflora, M. yukonensis
Subordinate taxa
M. nuttallii var. fragilis, M. nuttallii var. gracilis, M. nuttallii var. gregaria, M. nuttallii var. nuttallii
Synonyms Arenaria nuttallii, Minuopsis nuttallii Arenaria arctica, Alsine arctica, Lidia arctica
Name authority (Pax) Briquet: Annuaire Conserv. Jard. Bot. Genève 13–14: 385. (1911) (Steven ex Seringe) Graebner: in P. F. A. Ascherson et al., Syn. Mitteleur. Fl. 5(1): 772. (1918)
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