Minuartia douglasii |
Minuartia patula |
|
---|---|---|
Douglas' sandwort, Douglas' stitchwort |
Pitcher's stitchwort |
|
Habit | Plants annual. | Plants winter annual or annual. |
Taproots | filiform to somewhat thickened. |
filiform. |
Stems | erect to widely spreading, green or purple, 4–30 cm, stipitate-glandular distally, internodes of all stems 1–5 times as long as leaves. |
erect to ascending, green, 5–30 cm, glabrous or sometimes stipitate-glandular distally or throughout, internodes of all stems 1–7 times as long as leaves; wintering stems absent. |
Leaves | sometimes overlapping proximally, ± evenly spaced, connate proximally, with loose, scarious sheath 0.3–0.7 mm; blade straight to variously curved or coiled, green or purple, concave, 1–3 veined, linear, 5–40 × 0.2–0.4 mm, flexuous, margins not thickened, often scarious, sometimes ciliate or stipitate-glandular, apex green to purple, acute, navicular, dull, glabrous to stipitate-glandular; axillary leaves present proximally. |
overlapping proximally, connate proximally, with loose, scarious to herbaceous sheath 0.1–0.5 mm; blade straight to variously curved, green, flat, prominently 1-veined abaxially, linear, 2–20 × 0.5–1.5(–1.8) mm, flexuous, margins not thickened, herbaceous, smooth, apex green or purple, blunt to acute, flat, ± shiny, glabrous to stipitate-glandular; axillary leaves absent. |
Inflorescences | 7–15+-flowered, open cymes; bracts subulate, scarious. |
5–30-flowered, open cymes; bracts subulate to ovate, herbaceous. |
Pedicels | 0.2–4 cm, stipitate-glandular. |
0.3–3 cm, stipitate-glandular. |
Flowers | hypanthium disc-shaped; sepals 1(–3)-veined, midvein often more prominent than lateral veins, ± ovate (herbaceous portion lanceolate to oblong-lanceolate to narrowly ovate), 2.5–3.7 mm, not enlarging in fruit, apex often purple, obtuse to acute, not hooded, stipitate-glandular at least proximally; petals ovate, (1–)1.7–2.1 times as long as sepals, apex broadly rounded, entire or irregularly dentate. |
hypanthium shallowly disc-shaped; sepals prominently (3- or) 5-veined, narrowly to broadly lanceolate (herbaceous portion narrowly to broadly lanceolate), 4–5.5 mm, not enlarging in fruit, apex green or purple, narrowly acute to acuminate, not hooded, glabrous to sparsely stipitate-glandular; petals obovate, 1.5–2.2(–3) times as long as sepals, apex rounded, broadly notched. |
Capsules | on stipe 0.3–0.5 mm, broadly ovoid, 4 mm, longer than sepals. |
on stipe ca. 0.1 mm or shorter, narrowly ellipsoid, 3–4.2 mm, shorter than sepals. |
Seeds | reddish brown, winged, orbiculate with radicle not prolonged, compressed, 1.3–2 mm, tuberculate; tubercles low, elongate. |
reddish brown to black, suborbiculate, radicle obscure, slightly compressed, 0.5–0.6 mm, tuberculate; tubercles low, rounded. |
Minuartia douglasii |
Minuartia patula |
|
Phenology | Flowering spring–early summer. | Flowering spring–early summer. |
Habitat | Sandy and rocky slopes in chaparral, oak or pine woodlands | Prairies, meadows, limestone barrens, and rocky outcrops in sandy, clayey, or gravelly soils |
Elevation | 100-1800 m (300-5900 ft) | 0-500 |
Distribution |
AZ; CA; OR
|
AL; AR; GA; IL; IN; KS; KY; LA; MO; MS; OH; OK; PA; TN; TX; VA; WI |
Discussion | The winglike margins on the seeds of Minuartia douglasii are unique among North American members of the genus; M. howellii and M. tenella, the other members of sect. Greniera (Gay) Mattfeld, do not share this feature. Plants with petal apices usually emarginate rather than entire and obtuse and often shorter than in typical plants have been recognized by some as var. emarginata. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Minuartia patula and the related M. muscorum have received little attention in comparison to the granite-outcrop minuartias, the M. uniflora complex. J. A. Steyermark (1941) studied these taxa and described three forms, based chiefly on pubescence variation. Plants entirely glabrous [forma pitcheri (Nuttall) Steyermark] and those with sepals and pedicels somewhat stipitate-glandular (forma media Steyermark) were segregated from densely stipitate-glandular plants (forma patula). We do not feel that such variations deserve formal taxonomic recognition. Forma robusta, as defined by Steyermark, is here referred to M. muscorum. Most specimens of Minuartia patula have prominently five-veined sepals (seen especially easily in the glabrous forms); occasional plants from Georgia, Kentucky, and Virginia have glabrous sepals with only three strong veins, resembling those of M. muscorum; in other features, including the seeds, they are clearly referable to M. patula. The status of the plants with three-veined sepals remains ambiguous; J. A. Steyermark (1941) included them in his forma media and B. Maguire (1951) included them (in our opinion incorrectly) in his var. robusta. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 5, p. 124. | FNA vol. 5, p. 131. |
Parent taxa | Caryophyllaceae > subfam. Alsinoideae > Minuartia | Caryophyllaceae > subfam. Alsinoideae > Minuartia |
Sibling taxa | ||
Synonyms | Arenaria douglasii, M. douglasii var. emarginata | Arenaria patula, Alsinopsis patula, Alsinopsis pitcheri, Sabulina patula |
Name authority | (Fenzl ex Torrey & A. Gray) Mattfeld: Bot. Jahrb. Syst. 57(Beibl. 126): 27. (1921) | (Michaux) Mattfeld: Bot. Jahrb. Syst. 57(Beibl. 126): 28. (1921) |
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