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Saddle Mountain saxifrage

Howell's saxifrage

alpine saxifrage, Tolmie's alpine saxifrage, Tolmie's saxifrage

Habit Plants solitary or in clumps, short-rhizomatous. Plants solitary or in groups, with bulbils on caudices. Plants mat-forming, (stems trailing, thin, basally slightly woody), with slender, woody caudices.
Leaves

basal;

petiole often indistinct, flattened, 5–50 mm;

blade elliptic to obovate, 4–12 cm, slightly fleshy, base cuneate, margins serrate to dentate, densely ciliate, surfaces densely tangled-hairy.

basal;

petiole flattened, 1–4 cm;

blade oblong to narrowly ovate, 1–2(–3) cm, ± fleshy, base attenuate, margins sharply serrate, eciliate, surfaces sparsely tangled, reddish brown-hairy abaxially, glabrous adaxially.

cauline, proximally crowded;

petiole absent;

blade elliptic-obovate, 0.8–1.5 cm, fleshy, base cuneate, margins entire, sometimes proximally sparsely long-ciliate, surfaces glabrous.

Inflorescences

70+-flowered, open, lax, flat-topped thyrses, 15–35 cm, tangled, yellow-tipped stipitate-glandular.

5–10(–20)-flowered, very open, lax, ± flat-topped cymes or thyrses, 5–10(–20) cm, glabrate to sparsely purple-tipped stipitate-glandular; (bracts glabrous).

2–10-flowered, lax cymes, sometimes solitary flowers, 3–12 cm, glabrous or purplish- or pink-tipped stipitate-glandular; (bracts 1–3, proximal).

Flowers

sepals reflexed, elliptic to ovate;

petals white, not spotted, elliptic, clawed, 2–5 mm, longer than sepals;

filaments linear, flattened;

pistils connate to 1/2 their lengths;

ovary 1/2 inferior, appearing more superior in fruit.

sepals strongly reflexed, elliptic, (surfaces glabrous);

petals white, not spotted, obovate to elliptic, clawed, 2.5–4.5 mm, longer than sepals;

filaments linear, flattened;

pistils distinct almost to base;

ovary ± superior.

sepals erect to spreading, ovate;

petals white, not spotted, linear to oblanceolate, 2.5–5 mm, longer than sepals, short- or not clawed;

filaments broadly club-shaped (± petaloid);

pistils connate to 1/2 their lengths;

ovary ± superior (to 1/4 adnate to hypanthium).

Capsules

greenish to reddish purple, folliclelike.

purple, valvate.

green, purple mottled, valvate.

2n

= 76.

= 30.

Micranthes hitchcockiana

Micranthes howellii

Micranthes tolmiei

Phenology Flowering late spring–summer. Flowering early spring–early summer. Flowering summer.
Habitat Wet rocks and ledges on mountain-top balds Wet, rocky ledges and crevices Arctic and alpine tundra, fell-fields
Elevation 600-1100 m [2000-3600 ft] 0-900 m [0-3000 ft] 500-4000 m [1600-13100 ft]
Discussion

Of conservation concern.

Micranthes hitchcockiana may have originated from hybridization between M. rufidula and M. oregana (P. E. Elvander 1984; W. E. Perkins 1978).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Unlike those of most Micranthes species, the leaves of M. tolmiei are proximally cauline and the ovules have two integuments. The seeds have a loose, winglike testa.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Distribution
from FNA
OR
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[BONAP county map]
from FNA
CA; OR
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[BONAP county map]
from FNA
AK; CA; ID; MT; NV; OR; UT; WA; BC
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Parent taxa Saxifragaceae > Micranthes Saxifragaceae > Micranthes Saxifragaceae > Micranthes
Sibling taxa
M. apetala, M. aprica, M. bryophora, M. californica, M. calycina, M. careyana, M. caroliniana, M. eriophora, M. ferruginea, M. foliolosa, M. fragosa, M. gaspensis, M. gormanii, M. hieraciifolia, M. howellii, M. idahoensis, M. integrifolia, M. lyallii, M. marshallii, M. micranthidifolia, M. nelsoniana, M. nidifica, M. nivalis, M. nudicaulis, M. occidentalis, M. odontoloma, M. oregana, M. palmeri, M. pensylvanica, M. petiolaris, M. razshivinii, M. reflexa, M. rhomboidea, M. rufidula, M. spicata, M. stellaris, M. subapetala, M. tempestiva, M. tenuis, M. texana, M. tischii, M. tolmiei, M. unalaschcensis, M. virginiensis
M. apetala, M. aprica, M. bryophora, M. californica, M. calycina, M. careyana, M. caroliniana, M. eriophora, M. ferruginea, M. foliolosa, M. fragosa, M. gaspensis, M. gormanii, M. hieraciifolia, M. hitchcockiana, M. idahoensis, M. integrifolia, M. lyallii, M. marshallii, M. micranthidifolia, M. nelsoniana, M. nidifica, M. nivalis, M. nudicaulis, M. occidentalis, M. odontoloma, M. oregana, M. palmeri, M. pensylvanica, M. petiolaris, M. razshivinii, M. reflexa, M. rhomboidea, M. rufidula, M. spicata, M. stellaris, M. subapetala, M. tempestiva, M. tenuis, M. texana, M. tischii, M. tolmiei, M. unalaschcensis, M. virginiensis
M. apetala, M. aprica, M. bryophora, M. californica, M. calycina, M. careyana, M. caroliniana, M. eriophora, M. ferruginea, M. foliolosa, M. fragosa, M. gaspensis, M. gormanii, M. hieraciifolia, M. hitchcockiana, M. howellii, M. idahoensis, M. integrifolia, M. lyallii, M. marshallii, M. micranthidifolia, M. nelsoniana, M. nidifica, M. nivalis, M. nudicaulis, M. occidentalis, M. odontoloma, M. oregana, M. palmeri, M. pensylvanica, M. petiolaris, M. razshivinii, M. reflexa, M. rhomboidea, M. rufidula, M. spicata, M. stellaris, M. subapetala, M. tempestiva, M. tenuis, M. texana, M. tischii, M. unalaschcensis, M. virginiensis
Synonyms Saxifraga hitchcockiana Saxifraga howellii Saxifraga tolmiei, Saxifraga tolmiei var. ledifolia
Name authority (Elvander) Brouillet & Gornall: Novon 18: 138. 2008 , (Greene) Small: in N. L. Britton et al., N. Amer. Fl. 22: 140. (1905) (Torrey & A. Gray) Brouillet & Gornall: J. Bot. Res. Inst. Texas 1: 1022. 2007 ,
Source Flora of North America vol. 8, p. 70. Flora of North America vol. 8, p. 63. Flora of North America vol. 8, p. 54.
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