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San Joaquin blazing star

sand blazing star, white-bract blazing star

Habit Plants candelabra-form, (8–)20–50(–60) cm. Plants 7–35(–45) cm.
Basal leaves

persisting;

petiole present or absent;

blade lanceolate to linear, margins deeply to shallowly lobed.

petioles present or absent;

blade lanceolate, to 10(–16) cm;

margins shallowly to deeply lobed, lobes acute.

Cauline leaves

petiole absent;

blade ovate to linear, to 12 cm, margins deeply lobed to dentate.

petioles absent;

blade ovate to lanceolate, to 10(–16) cm, base often cordate-clasping, margins deeply to shallowly lobed, lobes acute.

Bracts

green, ovate to lanceolate, 6.6–12.8 × 1.9–6.8 mm, width 1/5–2/3 length, not concealing capsule, margins 3–7-lobed.

white with green margins, conspicuous, concealing pedicel, ovary, and capsule.

Flowers

sepals 3–13 mm;

petals red to orange proximally, orange to yellow distally, 8–22 mm, apex mucronate, rounded, or retuse;

stamens 20+, 4–11 mm, filaments monomorphic, filiform, unlobed;

styles 5–13 mm.

petals white to pale yellow, usually with orange veins, obovate, 13–65 mm, apex mucronate;

stamens 4–26 mm, less than 1/2 petal length, all fertile, none petaloid (flowers appearing to have 5 petals);

filaments ± monomorphic, linear, distally 2-lobed, lobes 0.5–2 mm;

anther stalk shorter or longer than filament lobes;

style 8–30 mm.

Capsules

clavate, 12–35 × 2–4 mm, axillary curved to 90° at maturity, usually inconspicuously longitudinally ribbed.

cylindric to cup-shaped, 14–25(–30) × 5–10 mm, erect.

Seeds

20–40, in 2+ rows distal to mid fruit, tan, dark-mottled or not, usually irregularly polygonal, occasionally triangular prisms proximal to mid fruit, surface tuberculate under 10x magnification; recurved flap over hilum absent;

seed coat cell outer periclinal wall domed, domes on seed edges more than 1/2 as tall as wide at maturity.

usually constricted proximal to middle;

seed coat anticlinal cell walls wavy.

2n

= 18.

= 18.

Mentzelia pectinata

Mentzelia involucrata

Phenology Flowering Mar–May. Flowering Jan–May.
Habitat Slopes of sandy or gray-white silty soils, grasslands, oak savannas, uncommonly juniper woodlands. Washes, fans, slopes, desert scrub.
Elevation 200–1400 m. (700–4600 ft.) 50–900 m. (200–3000 ft.)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; Mexico (Baja California, Sonora)
[WildflowerSearch map]
[BONAP county map]
Discussion

Mentzelia pectinata occurs in Kern, San Luis Obispo, Santa Barbara, and Tulare counties, around the southern rim of the San Joaquin Valley, extending into the Inner Coast Ranges and the southern foothills of the Sierra Nevada. Petal color varies from orange to yellow, and fully fertile artificial hybrids have been obtained between populations representing the extreme phenotypes (J. E. Zavortink 1966). Phylogenetic studies suggest that M. pectinata and M. congesta have hybridized to form several allopolyploid species (J. M. Brokaw and L. Hufford 2010b) despite their current allopatric distributions.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Mentzelia involucrata is easily distinguished from other species in sect. Bicuspidaria by its large white bracts, which are strikingly similar to those of M. congesta in sect. Trachyphytum. Phylogenetic studies show that these species are not closely related, suggesting a homoplasious origin of the characteristic (J. M. Brokaw and L. Hufford, 2010). Populations exhibiting petals longer than 3 cm have previously been called var. megalantha. However, a study by G. S. Daniels (1970) suggested that insufficient geographic and morphological discontinuities exist for recognition of varieties.

P. A. Munz (1974) mistakenly cited var. megalantha as Mentzelia involucrata subsp. megalantha I. M. Johnston, a name never validly published.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 541. FNA vol. 12, p. 525.
Parent taxa Loasaceae > Mentzelia > sect. Trachyphytum Loasaceae > Mentzelia > sect. Bicuspidaria
Sibling taxa
M. affinis, M. albescens, M. albicaulis, M. argillicola, M. argillosa, M. aspera, M. asperula, M. candelariae, M. canyonensis, M. chrysantha, M. collomiae, M. congesta, M. conspicua, M. crocea, M. cronquistii, M. decapetala, M. densa, M. desertorum, M. dispersa, M. eremophila, M. filifolia, M. floridana, M. flumensevera, M. goodrichii, M. gracilenta, M. hirsutissima, M. holmgreniorum, M. hualapaiensis, M. humilis, M. integra, M. involucrata, M. inyoensis, M. isolata, M. jonesii, M. laciniata, M. laevicaulis, M. lagarosa, M. leucophylla, M. librina, M. lindheimeri, M. lindleyi, M. longiloba, M. marginata, M. memorabilis, M. mexicana, M. micrantha, M. mollis, M. monoensis, M. montana, M. multicaulis, M. multiflora, M. nitens, M. nuda, M. obscura, M. oligosperma, M. oreophila, M. pachyrhiza, M. packardiae, M. paradoxensis, M. perennis, M. polita, M. procera, M. pterosperma, M. puberula, M. pumila, M. ravenii, M. reflexa, M. reverchonii, M. rhizomata, M. rusbyi, M. saxicola, M. shultziorum, M. sivinskii, M. speciosa, M. springeri, M. strictissima, M. thompsonii, M. tiehmii, M. todiltoensis, M. torreyi, M. tricuspis, M. tridentata, M. uintahensis, M. veatchiana
M. affinis, M. albescens, M. albicaulis, M. argillicola, M. argillosa, M. aspera, M. asperula, M. candelariae, M. canyonensis, M. chrysantha, M. collomiae, M. congesta, M. conspicua, M. crocea, M. cronquistii, M. decapetala, M. densa, M. desertorum, M. dispersa, M. eremophila, M. filifolia, M. floridana, M. flumensevera, M. goodrichii, M. gracilenta, M. hirsutissima, M. holmgreniorum, M. hualapaiensis, M. humilis, M. integra, M. inyoensis, M. isolata, M. jonesii, M. laciniata, M. laevicaulis, M. lagarosa, M. leucophylla, M. librina, M. lindheimeri, M. lindleyi, M. longiloba, M. marginata, M. memorabilis, M. mexicana, M. micrantha, M. mollis, M. monoensis, M. montana, M. multicaulis, M. multiflora, M. nitens, M. nuda, M. obscura, M. oligosperma, M. oreophila, M. pachyrhiza, M. packardiae, M. paradoxensis, M. pectinata, M. perennis, M. polita, M. procera, M. pterosperma, M. puberula, M. pumila, M. ravenii, M. reflexa, M. reverchonii, M. rhizomata, M. rusbyi, M. saxicola, M. shultziorum, M. sivinskii, M. speciosa, M. springeri, M. strictissima, M. thompsonii, M. tiehmii, M. todiltoensis, M. torreyi, M. tricuspis, M. tridentata, M. uintahensis, M. veatchiana
Synonyms M. involucrata var. megalantha
Name authority Kellogg: Proc. Calif. Acad. Sci. 3: 40, fig. 9. (1863) S. Watson: Proc. Amer. Acad. Arts 20: 367. (1885)
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