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San Joaquin blazing star

Arapien blazingstar

Habit Plants candelabra-form, (8–)20–50(–60) cm. Plants perennial, bushlike, with subterranean caudices.
Stems

multiple, erect, zigzag or straight;

branches distal or along entire stem, distal longest or all ± equal, antrorse, straight; hairy.

Leaves

blade 15.4–40(–86) × 4–12.2 mm, always some more than 10 mm wide, widest intersinus distance 4–12.2 mm;

proximal oblanceolate to spatulate, margins usually entire, rarely dentate, teeth 0(–4), perpendicular to leaf axis, 0.1–1 mm;

distal oblanceolate, spatulate, or elliptic, base not clasping, margins usually entire, rarely dentate, teeth 0(–4), perpendicular to leaf axis, 0.3–1.4 mm;

abaxial surface with simple grappling-hook, complex grappling-hook, and needlelike trichomes, adaxial surface with simple grappling-hook and needlelike trichomes.

Basal leaves

persisting;

petiole present or absent;

blade lanceolate to linear, margins deeply to shallowly lobed.

Cauline leaves

petiole absent;

blade ovate to linear, to 12 cm, margins deeply lobed to dentate.

Bracts

green, ovate to lanceolate, 6.6–12.8 × 1.9–6.8 mm, width 1/5–2/3 length, not concealing capsule, margins 3–7-lobed.

margins entire.

Flowers

sepals 3–13 mm;

petals red to orange proximally, orange to yellow distally, 8–22 mm, apex mucronate, rounded, or retuse;

stamens 20+, 4–11 mm, filaments monomorphic, filiform, unlobed;

styles 5–13 mm.

petals golden yellow, 8.2–12.2(–15.2) × 2.4–5 mm, apex acute, glabrous abaxially;

stamens golden yellow, 5 outermost petaloid, filaments narrowly spatulate, slightly clawed, 5.6–11(–13.5) × 1.4–3 mm, with anthers, second whorl with anthers;

anthers usually twisted after dehiscence, epidermis papillate;

styles 5.5–9.4 mm.

Capsules

clavate, 12–35 × 2–4 mm, axillary curved to 90° at maturity, usually inconspicuously longitudinally ribbed.

cup-shaped, 5–8.4 × 3.1–6 mm, base tapering to rounded, not longitudinally ridged.

Seeds

20–40, in 2+ rows distal to mid fruit, tan, dark-mottled or not, usually irregularly polygonal, occasionally triangular prisms proximal to mid fruit, surface tuberculate under 10x magnification; recurved flap over hilum absent;

seed coat cell outer periclinal wall domed, domes on seed edges more than 1/2 as tall as wide at maturity.

coat anticlinal cell walls straight, papillae 3–5 per cell.

2n

= 18.

= 22.

Mentzelia pectinata

Mentzelia argillosa

Phenology Flowering Mar–May. Flowering Jun–Aug.
Habitat Slopes of sandy or gray-white silty soils, grasslands, oak savannas, uncommonly juniper woodlands. Sparsely vegetated steep cliffs or slopes composed of gypsum-rich clayey and gravelly soils.
Elevation 200–1400 m. (700–4600 ft.) 1600–1900 m. (5200–6200 ft.)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
UT
[BONAP county map]
Discussion

Mentzelia pectinata occurs in Kern, San Luis Obispo, Santa Barbara, and Tulare counties, around the southern rim of the San Joaquin Valley, extending into the Inner Coast Ranges and the southern foothills of the Sierra Nevada. Petal color varies from orange to yellow, and fully fertile artificial hybrids have been obtained between populations representing the extreme phenotypes (J. E. Zavortink 1966). Phylogenetic studies suggest that M. pectinata and M. congesta have hybridized to form several allopolyploid species (J. M. Brokaw and L. Hufford 2010b) despite their current allopatric distributions.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Mentzelia argillosa is endemic to the Arapien Shale formation in Sevier and Sanpete counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 541. FNA vol. 12, p. 507.
Parent taxa Loasaceae > Mentzelia > sect. Trachyphytum Loasaceae > Mentzelia > sect. Bartonia
Sibling taxa
M. affinis, M. albescens, M. albicaulis, M. argillicola, M. argillosa, M. aspera, M. asperula, M. candelariae, M. canyonensis, M. chrysantha, M. collomiae, M. congesta, M. conspicua, M. crocea, M. cronquistii, M. decapetala, M. densa, M. desertorum, M. dispersa, M. eremophila, M. filifolia, M. floridana, M. flumensevera, M. goodrichii, M. gracilenta, M. hirsutissima, M. holmgreniorum, M. hualapaiensis, M. humilis, M. integra, M. involucrata, M. inyoensis, M. isolata, M. jonesii, M. laciniata, M. laevicaulis, M. lagarosa, M. leucophylla, M. librina, M. lindheimeri, M. lindleyi, M. longiloba, M. marginata, M. memorabilis, M. mexicana, M. micrantha, M. mollis, M. monoensis, M. montana, M. multicaulis, M. multiflora, M. nitens, M. nuda, M. obscura, M. oligosperma, M. oreophila, M. pachyrhiza, M. packardiae, M. paradoxensis, M. perennis, M. polita, M. procera, M. pterosperma, M. puberula, M. pumila, M. ravenii, M. reflexa, M. reverchonii, M. rhizomata, M. rusbyi, M. saxicola, M. shultziorum, M. sivinskii, M. speciosa, M. springeri, M. strictissima, M. thompsonii, M. tiehmii, M. todiltoensis, M. torreyi, M. tricuspis, M. tridentata, M. uintahensis, M. veatchiana
M. affinis, M. albescens, M. albicaulis, M. argillicola, M. aspera, M. asperula, M. candelariae, M. canyonensis, M. chrysantha, M. collomiae, M. congesta, M. conspicua, M. crocea, M. cronquistii, M. decapetala, M. densa, M. desertorum, M. dispersa, M. eremophila, M. filifolia, M. floridana, M. flumensevera, M. goodrichii, M. gracilenta, M. hirsutissima, M. holmgreniorum, M. hualapaiensis, M. humilis, M. integra, M. involucrata, M. inyoensis, M. isolata, M. jonesii, M. laciniata, M. laevicaulis, M. lagarosa, M. leucophylla, M. librina, M. lindheimeri, M. lindleyi, M. longiloba, M. marginata, M. memorabilis, M. mexicana, M. micrantha, M. mollis, M. monoensis, M. montana, M. multicaulis, M. multiflora, M. nitens, M. nuda, M. obscura, M. oligosperma, M. oreophila, M. pachyrhiza, M. packardiae, M. paradoxensis, M. pectinata, M. perennis, M. polita, M. procera, M. pterosperma, M. puberula, M. pumila, M. ravenii, M. reflexa, M. reverchonii, M. rhizomata, M. rusbyi, M. saxicola, M. shultziorum, M. sivinskii, M. speciosa, M. springeri, M. strictissima, M. thompsonii, M. tiehmii, M. todiltoensis, M. torreyi, M. tricuspis, M. tridentata, M. uintahensis, M. veatchiana
Synonyms Nuttallia argillosa
Name authority Kellogg: Proc. Calif. Acad. Sci. 3: 40, fig. 9. (1863) J. Darlington: Ann. Missouri Bot. Gard. 21: 153. (1934)
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