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San Joaquin blazing star

Pioche blazingstar, Pioche stickleaf

Habit Plants candelabra-form, (8–)20–50(–60) cm. Plants perennial, bushlike, with subterranean caudices.
Stems

multiple, erect or decumbent, straight;

branches along entire stem, distal longest, antrorse, straight; hairy.

Leaves

blade (13.5–)16.4–33.3 × 1.5–8.8 mm, widest intersinus distance 1.5–6.1 mm;

proximal oblanceolate to spatulate, margins entire or pinnate, lobes 0–8 (always some leaves with 5+ lobes), perpendicular to leaf axis, 0.5–2 mm, always some more than 1.4 mm;

distal oblanceolate, elliptic, lanceolate, or linear, base not clasping, margins entire, dentate, or pinnate, teeth or lobes 0–6, perpendicular to leaf axis, 0.1–2 mm;

abaxial surface with complex grappling-hook, needlelike, and occasionally simple grappling-hook trichomes, adaxial surface with needlelike and generally simple grappling-hook trichomes.

Basal leaves

persisting;

petiole present or absent;

blade lanceolate to linear, margins deeply to shallowly lobed.

Cauline leaves

petiole absent;

blade ovate to linear, to 12 cm, margins deeply lobed to dentate.

Bracts

green, ovate to lanceolate, 6.6–12.8 × 1.9–6.8 mm, width 1/5–2/3 length, not concealing capsule, margins 3–7-lobed.

margins entire.

Flowers

sepals 3–13 mm;

petals red to orange proximally, orange to yellow distally, 8–22 mm, apex mucronate, rounded, or retuse;

stamens 20+, 4–11 mm, filaments monomorphic, filiform, unlobed;

styles 5–13 mm.

petals golden yellow, 7.2–11.9 × 2.7–4.9 mm, apex acute to rounded, glabrous abaxially;

stamens golden yellow, 5 outermost petaloid, filaments narrowly spatulate, slightly clawed, 5.5–10.3 × 1.4–2.7 mm, with anthers, second whorl with anthers;

anthers usually twisted after dehiscence, epidermis papillate;

styles 5–9 mm.

Capsules

clavate, 12–35 × 2–4 mm, axillary curved to 90° at maturity, usually inconspicuously longitudinally ribbed.

cup-shaped, 4.9–7.6 × 3.4–5.5 mm, base rounded to tapering, not longitudinally ridged.

Seeds

20–40, in 2+ rows distal to mid fruit, tan, dark-mottled or not, usually irregularly polygonal, occasionally triangular prisms proximal to mid fruit, surface tuberculate under 10x magnification; recurved flap over hilum absent;

seed coat cell outer periclinal wall domed, domes on seed edges more than 1/2 as tall as wide at maturity.

coat anticlinal cell walls straight, papillae 5–8 per cell.

2n

= 18.

= 22.

Mentzelia pectinata

Mentzelia argillicola

Phenology Flowering Mar–May. Flowering Jun–Aug.
Habitat Slopes of sandy or gray-white silty soils, grasslands, oak savannas, uncommonly juniper woodlands. Sparsely vegetated cliffs and knolls, gypsum-rich clayey soils.
Elevation 200–1400 m. (700–4600 ft.) 1400–1900 m. (4600–6200 ft.)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
NV
[BONAP county map]
Discussion

Mentzelia pectinata occurs in Kern, San Luis Obispo, Santa Barbara, and Tulare counties, around the southern rim of the San Joaquin Valley, extending into the Inner Coast Ranges and the southern foothills of the Sierra Nevada. Petal color varies from orange to yellow, and fully fertile artificial hybrids have been obtained between populations representing the extreme phenotypes (J. E. Zavortink 1966). Phylogenetic studies suggest that M. pectinata and M. congesta have hybridized to form several allopolyploid species (J. M. Brokaw and L. Hufford 2010b) despite their current allopatric distributions.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Mentzelia argillicola is known only from Lincoln County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 541. FNA vol. 12, p. 507.
Parent taxa Loasaceae > Mentzelia > sect. Trachyphytum Loasaceae > Mentzelia > sect. Bartonia
Sibling taxa
M. affinis, M. albescens, M. albicaulis, M. argillicola, M. argillosa, M. aspera, M. asperula, M. candelariae, M. canyonensis, M. chrysantha, M. collomiae, M. congesta, M. conspicua, M. crocea, M. cronquistii, M. decapetala, M. densa, M. desertorum, M. dispersa, M. eremophila, M. filifolia, M. floridana, M. flumensevera, M. goodrichii, M. gracilenta, M. hirsutissima, M. holmgreniorum, M. hualapaiensis, M. humilis, M. integra, M. involucrata, M. inyoensis, M. isolata, M. jonesii, M. laciniata, M. laevicaulis, M. lagarosa, M. leucophylla, M. librina, M. lindheimeri, M. lindleyi, M. longiloba, M. marginata, M. memorabilis, M. mexicana, M. micrantha, M. mollis, M. monoensis, M. montana, M. multicaulis, M. multiflora, M. nitens, M. nuda, M. obscura, M. oligosperma, M. oreophila, M. pachyrhiza, M. packardiae, M. paradoxensis, M. perennis, M. polita, M. procera, M. pterosperma, M. puberula, M. pumila, M. ravenii, M. reflexa, M. reverchonii, M. rhizomata, M. rusbyi, M. saxicola, M. shultziorum, M. sivinskii, M. speciosa, M. springeri, M. strictissima, M. thompsonii, M. tiehmii, M. todiltoensis, M. torreyi, M. tricuspis, M. tridentata, M. uintahensis, M. veatchiana
M. affinis, M. albescens, M. albicaulis, M. argillosa, M. aspera, M. asperula, M. candelariae, M. canyonensis, M. chrysantha, M. collomiae, M. congesta, M. conspicua, M. crocea, M. cronquistii, M. decapetala, M. densa, M. desertorum, M. dispersa, M. eremophila, M. filifolia, M. floridana, M. flumensevera, M. goodrichii, M. gracilenta, M. hirsutissima, M. holmgreniorum, M. hualapaiensis, M. humilis, M. integra, M. involucrata, M. inyoensis, M. isolata, M. jonesii, M. laciniata, M. laevicaulis, M. lagarosa, M. leucophylla, M. librina, M. lindheimeri, M. lindleyi, M. longiloba, M. marginata, M. memorabilis, M. mexicana, M. micrantha, M. mollis, M. monoensis, M. montana, M. multicaulis, M. multiflora, M. nitens, M. nuda, M. obscura, M. oligosperma, M. oreophila, M. pachyrhiza, M. packardiae, M. paradoxensis, M. pectinata, M. perennis, M. polita, M. procera, M. pterosperma, M. puberula, M. pumila, M. ravenii, M. reflexa, M. reverchonii, M. rhizomata, M. rusbyi, M. saxicola, M. shultziorum, M. sivinskii, M. speciosa, M. springeri, M. strictissima, M. thompsonii, M. tiehmii, M. todiltoensis, M. torreyi, M. tricuspis, M. tridentata, M. uintahensis, M. veatchiana
Name authority Kellogg: Proc. Calif. Acad. Sci. 3: 40, fig. 9. (1863) N. H. Holmgren & P. K. Holmgren: Syst. Bot. 27: 751, fig. 3. (2002)
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