Mentzelia humilis
Mentzelia pectinata
gypsum blazingstar
San Joaquin blazing star
solitary or multiple, erect, straight;
branches distal or along entire stem, distal longest or all ± equal, antrorse; straight to upcurved; hairy.
blade 25–95 × 5.5–28(–36.2) mm, widest intersinus distance 0.8–9.1 mm;
proximal spatulate to oblanceolate or elliptic, margins pinnate to pinnatisect, lobes 4–16(–20), slightly antrorse or perpendicular to leaf axis, 2.3–11.8(–16.4) mm;
distal elliptic, lanceolate, spatulate, or linear, base not clasping, margins entire or dentate to pinnatisect, teeth or lobes (0–)4–16, slightly antrorse or perpendicular to leaf axis, 2.3–13.8(–16.9) mm;
abaxial surface with simple grappling-hook, needlelike, and sometimes complex grappling-hook trichomes, adaxial surface with simple grappling-hook and needlelike trichomes.
persisting;
petiole present or absent;
blade lanceolate to linear, margins deeply to shallowly lobed.
petiole absent;
blade ovate to linear, to 12 cm, margins deeply lobed to dentate.
margins usually entire, rarely pinnate.
green, ovate to lanceolate, 6.6–12.8 × 1.9–6.8 mm, width 1/5–2/3 length, not concealing capsule, margins 3–7-lobed.
petals white, 10.3–13(–28.6) × 1.4–4 mm, apex acute, glabrous abaxially;
stamens white, 5 outermost petaloid, filaments linear to narrowly spatulate, slightly clawed, 8.8–19(–22.3) × 0.7–3.3 mm, without anthers, second whorl without anthers;
anthers straight after dehiscence, epidermis papillate;
styles 6.5–11.3 mm.
sepals 3–13 mm;
petals red to orange proximally, orange to yellow distally, 8–22 mm, apex mucronate, rounded, or retuse;
stamens 20+, 4–11 mm, filaments monomorphic, filiform, unlobed;
styles 5–13 mm.
cup-shaped, 5.3–10.2 × (4.2–)5.2–8.6 mm, base rounded, not longitudinally ridged.
clavate, 12–35 × 2–4 mm, axillary curved to 90° at maturity, usually inconspicuously longitudinally ribbed.
coat anticlinal cell walls wavy, papillae 6–12 per cell.
20–40, in 2+ rows distal to mid fruit, tan, dark-mottled or not, usually irregularly polygonal, occasionally triangular prisms proximal to mid fruit, surface tuberculate under 10x magnification; recurved flap over hilum absent;
seed coat cell outer periclinal wall domed, domes on seed edges more than 1/2 as tall as wide at maturity.
= 18.
Mentzelia humilis
Mentzelia pectinata
Varieties 2 (2 in the flora).
The basionym of Mentzelia humilis is often cited as M. multiflora var. humilis A. Gray (1852). However, in that publication Gray indicated accepted names in Roman capitals (see ipni.org), whereas the varietal name is in lower case, indicating that he was using it as a descriptive term rather than a scientific name. Valid publication of the basionym thus must be attributed to Urban and Gilg.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Mentzelia pectinata occurs in Kern, San Luis Obispo, Santa Barbara, and Tulare counties, around the southern rim of the San Joaquin Valley, extending into the Inner Coast Ranges and the southern foothills of the Sierra Nevada. Petal color varies from orange to yellow, and fully fertile artificial hybrids have been obtained between populations representing the extreme phenotypes (J. E. Zavortink 1966). Phylogenetic studies suggest that M. pectinata and M. congesta have hybridized to form several allopolyploid species (J. M. Brokaw and L. Hufford 2010b) despite their current allopatric distributions.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Leaf blade widest intersinus distances 0.8–2.6(–3.3) mm; lobes of proximal leaves 1–2.9 mm wide. | var. humilis |
1. Leaf blade widest intersinus distances 1.6–9.1 mm (always on some leaves greater than 2.6 mm); lobes of proximal leaves (1.1–)2.9–5.1 mm wide. | var. guadalupensis |
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