Mentzelia humilis |
Mentzelia pachyrhiza |
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gypsum blazingstar |
big-root stickleaf, Coahuila blazingstar |
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Habit | Plants perennial, bushlike, with ground-level caudices. | Plants perennial, with tubers. | ||||
Stems | solitary or multiple, erect, straight; branches distal or along entire stem, distal longest or all ± equal, antrorse; straight to upcurved; hairy. |
erect, to 50 cm. |
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Leaves | blade 25–95 × 5.5–28(–36.2) mm, widest intersinus distance 0.8–9.1 mm; proximal spatulate to oblanceolate or elliptic, margins pinnate to pinnatisect, lobes 4–16(–20), slightly antrorse or perpendicular to leaf axis, 2.3–11.8(–16.4) mm; distal elliptic, lanceolate, spatulate, or linear, base not clasping, margins entire or dentate to pinnatisect, teeth or lobes (0–)4–16, slightly antrorse or perpendicular to leaf axis, 2.3–13.8(–16.9) mm; abaxial surface with simple grappling-hook, needlelike, and sometimes complex grappling-hook trichomes, adaxial surface with simple grappling-hook and needlelike trichomes. |
petiole mostly 3–20 mm, less than 3 mm only on smallest, distalmost leaves; blade ovate to hastate, basally lobed or unlobed, to 4 × 3 cm, base usually acute to obtuse, sometimes attenuate or hastate, margins usually serrate, sometimes crenate, apex acute. |
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Bracts | margins usually entire, rarely pinnate. |
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Pedicels | (fruiting) (0–)2 × 2 mm. |
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Flowers | petals white, 10.3–13(–28.6) × 1.4–4 mm, apex acute, glabrous abaxially; stamens white, 5 outermost petaloid, filaments linear to narrowly spatulate, slightly clawed, 8.8–19(–22.3) × 0.7–3.3 mm, without anthers, second whorl without anthers; anthers straight after dehiscence, epidermis papillate; styles 6.5–11.3 mm. |
petals orange, 5–11.5 × 3–5 mm, apex acute to rounded, hairy abaxially on distal 1/2; stamens 15–45, 4–8 mm, filaments usually monomorphic, filiform, rarely heteromorphic, outermost slightly spatulate, inner filiform; style 3–7.5 mm. |
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Capsules | cup-shaped, 5.3–10.2 × (4.2–)5.2–8.6 mm, base rounded, not longitudinally ridged. |
subcylindric to clavate, 6–10 × 1.5–2.3 mm, base tapering gradually, capsule and pedicel not well-differentiated, walls thin, brittle. |
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Seeds | coat anticlinal cell walls wavy, papillae 6–12 per cell. |
2–3 per capsule, oblong, with transverse folds. |
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2n | = 22. |
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Mentzelia humilis |
Mentzelia pachyrhiza |
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Phenology | Flowering May–Nov. | |||||
Habitat | Steep limestone cliffs, gravelly slopes of gypseous clayey soils, desert scrub and Larrea communities. | |||||
Elevation | 900–2000 m. (3000–6600 ft.) | |||||
Distribution |
NM; TX
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TX; Mexico (Chihuahua, Coahuila, Durango, Nuevo León) |
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Discussion | Varieties 2 (2 in the flora). The basionym of Mentzelia humilis is often cited as M. multiflora var. humilis A. Gray (1852). However, in that publication Gray indicated accepted names in Roman capitals (see ipni.org), whereas the varietal name is in lower case, indicating that he was using it as a descriptive term rather than a scientific name. Valid publication of the basionym thus must be attributed to Urban and Gilg. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Mentzelia pachyrhiza, native to the Chihuahuan Desert, reaches the flora area only in southern Brewster and Presidio counties. In southwestern Texas and northeastern Chihuahua, where M. pachyrhiza and M. oligosperma have overlapping ranges, H. J. Thompson and A. M. Powell (1981) reported that the former was found at elevations usually below 1100 meters in the Larrea zone, below the elevation of junipers and M. oligosperma. Thompson and Powell allied M. pachyrhiza with M. oligosperma and the South American M. grisebachii Urban & Gilg, now treated as a synonym of M. parvifolia Urban & Gilg ex Kurtz (M. Weigend 2007b). This placement is consistent with phylogenetic studies (J. Grissom and L. Hufford, unpubl.), which show that M. pachyrhiza is the sister species of M. parvifolia; they together are the sister of M. oligosperma and the Mexican M. pattersonii B. L. Turner. I. M. Johnston’s (1940) assertion that the large tuber of M. pachyrhiza is unique in sect. Mentzelia is incorrect. Tubers, often carrot-shaped, are common among the perennial species of the section. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 12, p. 503. | FNA vol. 12, p. 529. | ||||
Parent taxa | Loasaceae > Mentzelia > sect. Bartonia | Loasaceae > Mentzelia > sect. Mentzelia | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | M. pumila var. humilis, Nuttallia humilis, Touterea humilis | |||||
Name authority | (Urban & Gilg) J. Darlington: Ann. Missouri Bot. Gard. 21: 155. (1934) | I. M. Johnston: J. Arnold Arbor. 21: 71. (1940) | ||||
Web links |