Melica smithii |
Melica |
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mélique de Smith, Smith oniongrass, Smith's melic, Smith's melic grass, Smith's oniongrass |
melic, melicgrass, oniongrass |
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Habit | Plants loosely cespitose, not rhizomatous. | Plants perennial; cespitose or soboliferous, not or only shortly rhizomatous. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Culms | 60-160 cm, thickened basally, sometimes appearing cormous; internodes sometimes pubescent below the nodes. |
(4)9-250 cm, sometimes forming a basal corm; nodes and internodes usually glabrous. |
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Sheaths | usually glabrous, sometimes pilose or retrorsely scabrous, particularly at the throat, veins often prominent; ligules 2-4 mm; blades 15-25 cm long, 5-12 mm wide, both surfaces usually scabridulous, glabrous, sometimes the adaxial surfaces with hairs. |
closed almost to the top; auricles sometimes present; ligules thinly membranous, erose to lacerate, usually glabrous, those of the lower leaves shorter than those of the upper leaves; blades flat or folded, glabrous or hairy, particularly on the adaxial surfaces, sometimes scabrous. |
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Panicles | 12-30 cm; branches 7-11 cm, spreading to reflexed, with 4-7 spikelets, spikelets restricted to the distal portion, axils frequently with brownish pulvini; pedicels straight; disarticulation above the glumes. |
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Inflorescences | terminal panicles; primary branches often appressed; secondary branches appressed or divergent; pedicels either more or less straight or sharply bent below the spikelets, scabrous to strigose distally; disarticulation below the glumes in species with sharply bent pedicels, above the glumes in other species. |
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Spikelets | 12-18 mm, with 3-5 bisexual florets; rachilla internodes 2.5-3 mm. |
with 1-7 bisexual florets, terminating in a sterile structure, the rudiment, composed of 1-4 sterile florets; rudiments sometimes morphologically distinct from the bisexual florets, sometimes similar but smaller. |
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Glumes | membranous or chartaceous, distal margins wide, translucent; lower glumes 1-9-veined; upper glumes 1-11-veined; calluses glabrous; lemmas membranous basally, sometimes becoming coriaceous at maturity, glabrous or with hairs, (4)5-15-veined, usually unawned, sometimes awned, awns to 12 mm, straight; paleas from 1/2 as long as to almost equaling the lemmas, keels usually ciliate; lodicules fused into a single, collarlike structure extending 1/2 - 2/3 around the base of the ovaries; anthers (2)3. |
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Lower glumes | 4.5-7 mm long, 1-1.5 mm wide, 1-3-veined; upper glumes 6.5-9 mm long, 1.2-1.8 mm wide, 3-5-veined; lemmas 9.5-12 mm, glabrous or scabrous, 7-veined, apices bifid to emarginate, awned, awns 3-10 mm; paleas about 2/3 the length of the lemmas; anthers 1.3-2.5 mm; rudiments 3.5-6 mm, tapering, resembling the bisexual florets. |
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Caryopses | usually 2-3 mm, smooth, glabrous, longitudinally furrowed, falling from the floret when mature, x = 9. |
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2n | = unknown. |
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Melica smithii |
Melica |
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Distribution |
ID; MI; MT; OR; SD; WA; WI; WY; AB; BC; ON; QC
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AK; AL; AR; AZ; CA; CO; DC; FL; GA; IA; ID; IL; IN; KS; KY; LA; MD; MI; MN; MO; MS; MT; NC; NE; NJ; NM; NV; NY; OH; OK; OR; PA; SC; SD; TN; TX; UT; VA; WA; WI; WV; WY; AB; BC; ON; QC; SK |
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Discussion | Melica smithii grows in cool, moist woods from British Columbia and Alberta south to Oregon and Wyoming and, as a disjunct, from the Great Lakes region to western Quebec. It often forms colonies in the eastern portion of its range. Its disjunct distribution pattern is unusual among North America's grasses. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Melica includes approximately 80 species, which grow in all temperate regions of the world except Australia, usually in shady woodlands on dry stony slopes (Mejia-Saules and Bisby 2003). The species are relatively nutritious, but are rarely sufficiently abundant to be important as forage. Nineteen species of Melica grow in the Flora region. Two European species are grown as ornamentals in North America. Many of the seventeen native species merit such use. Several proposals have been made for dividing Melica into smaller units. American taxonomists have tended to favor Thurber's (1880) recognition of two subgenera: Melica and Bromelica. In subg. Melica, the pedicels are straight and disarticulation is above the glumes; in subg. Bromelica, the pedicels are sharply bent and the spikelets disarticulate below the glumes. Hempel (1970) recognized three subgenera in Melica, but his groups do not correspond well to the pattern of morphological variation seen in North America. More recently, Mejia-Saules and Bisby (2003) examined the variation in lemma silica bodies and hooked papillae within Melica. Their results are not consistent with either Thurber's or Hempel's treatment, but provide some support for Papp's (1928) recognition of two groups, based on the presence or absence of hairs on the lemmas and the compression of the spikelets. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 24, p. 95. | FNA vol. 24, p. 88. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Name authority | (Porter ex A. Gray) Vasey | L. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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