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mélique de Smith, Smith oniongrass, Smith's melic, Smith's melic grass, Smith's oniongrass

melic, melicgrass, oniongrass

Habit Plants loosely cespitose, not rhizomatous. Plants perennial; cespitose or soboliferous, not or only shortly rhizomatous.
Culms

60-160 cm, thickened basally, sometimes appearing cormous;

internodes sometimes pubescent below the nodes.

(4)9-250 cm, sometimes forming a basal corm;

nodes and internodes usually glabrous.

Sheaths

usually glabrous, sometimes pilose or retrorsely scabrous, particularly at the throat, veins often prominent;

ligules 2-4 mm;

blades 15-25 cm long, 5-12 mm wide, both surfaces usually scabridulous, glabrous, sometimes the adaxial surfaces with hairs.

closed almost to the top;

auricles sometimes present;

ligules thinly membranous, erose to lacerate, usually glabrous, those of the lower leaves shorter than those of the upper leaves;

blades flat or folded, glabrous or hairy, particularly on the adaxial surfaces, sometimes scabrous.

Panicles

12-30 cm;

branches 7-11 cm, spreading to reflexed, with 4-7 spikelets, spikelets restricted to the distal portion, axils frequently with brownish pulvini;

pedicels straight;

disarticulation above the glumes.

Inflorescences

terminal panicles;

primary branches often appressed;

secondary branches appressed or divergent;

pedicels either more or less straight or sharply bent below the spikelets, scabrous to strigose distally;

disarticulation below the glumes in species with sharply bent pedicels, above the glumes in other species.

Spikelets

12-18 mm, with 3-5 bisexual florets;

rachilla internodes 2.5-3 mm.

with 1-7 bisexual florets, terminating in a sterile structure, the rudiment, composed of 1-4 sterile florets;

rudiments sometimes morphologically distinct from the bisexual florets, sometimes similar but smaller.

Glumes

membranous or chartaceous, distal margins wide, translucent;

lower glumes 1-9-veined;

upper glumes 1-11-veined;

calluses glabrous;

lemmas membranous basally, sometimes becoming coriaceous at maturity, glabrous or with hairs, (4)5-15-veined, usually unawned, sometimes awned, awns to 12 mm, straight;

paleas from 1/2 as long as to almost equaling the lemmas, keels usually ciliate;

lodicules fused into a single, collarlike structure extending 1/2 - 2/3 around the base of the ovaries;

anthers (2)3.

Lower glumes

4.5-7 mm long, 1-1.5 mm wide, 1-3-veined;

upper glumes 6.5-9 mm long, 1.2-1.8 mm wide, 3-5-veined;

lemmas 9.5-12 mm, glabrous or scabrous, 7-veined, apices bifid to emarginate, awned, awns 3-10 mm;

paleas about 2/3 the length of the lemmas;

anthers 1.3-2.5 mm;

rudiments 3.5-6 mm, tapering, resembling the bisexual florets.

Caryopses

usually 2-3 mm, smooth, glabrous, longitudinally furrowed, falling from the floret when mature, x = 9.

2n

= unknown.

Melica smithii

Melica

Distribution
from FNA
ID; MI; MT; OR; SD; WA; WI; WY; AB; BC; ON; QC
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AL; AR; AZ; CA; CO; DC; FL; GA; IA; ID; IL; IN; KS; KY; LA; MD; MI; MN; MO; MS; MT; NC; NE; NJ; NM; NV; NY; OH; OK; OR; PA; SC; SD; TN; TX; UT; VA; WA; WI; WV; WY; AB; BC; ON; QC; SK
[BONAP county map]
Discussion

Melica smithii grows in cool, moist woods from British Columbia and Alberta south to Oregon and Wyoming and, as a disjunct, from the Great Lakes region to western Quebec. It often forms colonies in the eastern portion of its range. Its disjunct distribution pattern is unusual among North America's grasses.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Melica includes approximately 80 species, which grow in all temperate regions of the world except Australia, usually in shady woodlands on dry stony slopes (Mejia-Saules and Bisby 2003). The species are relatively nutritious, but are rarely sufficiently abundant to be important as forage.

Nineteen species of Melica grow in the Flora region. Two European species are grown as ornamentals in North America. Many of the seventeen native species merit such use.

Several proposals have been made for dividing Melica into smaller units. American taxonomists have tended to favor Thurber's (1880) recognition of two subgenera: Melica and Bromelica. In subg. Melica, the pedicels are straight and disarticulation is above the glumes; in subg. Bromelica, the pedicels are sharply bent and the spikelets disarticulate below the glumes. Hempel (1970) recognized three subgenera in Melica, but his groups do not correspond well to the pattern of morphological variation seen in North America. More recently, Mejia-Saules and Bisby (2003) examined the variation in lemma silica bodies and hooked papillae within Melica. Their results are not consistent with either Thurber's or Hempel's treatment, but provide some support for Papp's (1928) recognition of two groups, based on the presence or absence of hairs on the lemmas and the compression of the spikelets.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Spikelets disarticulating below the glumes; pedicels sharply bent just below the spikelets.
→ 2
2. Lemmas with hairs.
→ 3
3. Lemmas with hairs on the lower portion of the lemmas, the hairs twisted
M. montezumae
3. Lemmas with hairs on the marginal veins, the hairs not twisted
M. ciliata
2. Lemmas glabrous, sometimes scabridulous to scabrous.
→ 4
4. Rudiments acute to acuminate, similar to but smaller than the bisexual florets.
→ 5
5. Spikelets broadly V-shaped when mature, 5-13 mm wide; upper glumes 6-18 mm long
M. stricta
5. Spikelets parallel-sided when mature, 1.5-5 mm wide; upper glumes 5-8 mm long
M. porteri
4. Rudiments clublike, not resembling the bisexual florets.
→ 6
6. Rudiments at an angle to the rachilla; panicle branches with 2-5 spikelets
M. mutica
6. Rudiments in a straight line with the rachilla; panicle branches with 5-20 spikelets.
→ 7
7. Panicle branches often divergent to reflexed; glumes unequal, lower glumes shorter and more ovate than the upper glumes
M. nitens
7. Panicle branches strongly ascending to appressed; glumes subequal in length and similar in shape
M. altissima
1. Spikelets disarticulating above the glumes; pedicels more or less straight.
→ 8
8. Rudiments truncate to acute, not resembling the lowest florets.
→ 9
9. Bisexual florets 1(2); paleas almost as long as the lemmas.
→ 10
10. Rudiments shorter than the terminal rachilla internode; bisexual lemmas scabridulous, sometimes hairy
M. torreyana
10. Rudiments longer than the terminal rachilla internode; bisexual lemmas glabrous, sometimes scabrous
M. imperfecta
9. Bisexual florets 2-7; paleas 1/2 – 3/4 the length of the lemmas.
→ 11
11. Culm bases not forming distinct corms
M. californica
11. Culm bases forming distinct corms.
→ 12
12. Glumes usually less than 1/2 as long as the spikelets; ligules 0.1-2 mm long; corms connected to the rhizomes by a rootlike structure
M. spectabilis
12. Glumes from 1/2(2/3) as long as to equaling the spikelets; ligules 2-6 mm long; corms almost sessile on the rhizomes
M. bulbosa
8. Rudiments tapering, smaller than but otherwise similar to the lowest florets in shape.
→ 13
13. Lemmas awned.
→ 14
14. Awns shorter than 3 mm.
→ 15
15. Panicle branches appressed; lemmas usually with 0.7-1.3 mm hairs on the margins
M. harfordii
15. Panicle branches widespread to reflexed; lemmas glabrous
M. geyeri
14. Awns 3-12 mm long.
→ 16
16. Panicle branches 4-6 cm long, appressed or ascending; blades 2-6 mm wide
M. aristata
16. Panicle branches 7-11 cm long, spreading to reflexed; blades 5-12 mm wide
M. smithii
13. Lemmas unawned.
→ 17
17. Lemmas strongly tapering and acuminate, the veins usually hairy
M. subulata
17. Lemmas acute to obtuse, the veins hairy or not.
→ 18
18. Lemmas pubescent, the hairs on the marginal veins clearly longer than the hairs elsewhere
M. harfordii
18. Lemmas glabrous, scabrous, or pubescent, never with clearly longer hairs on the marginal veins.
→ 19
19. Rachilla internodes swollen when fresh, wrinkled when dry
M. fugax
19. Rachilla internodes not swollen when fresh, not wrinkled when dry.
→ 20
20. Panicle branches with 5-15 spikelets; paleas about 1/2 as long as the lemmas; culms not forming corms
M. frutescens
20. Panicle branches with 1-6 spikelets; paleas from 2/3 as long as to equaling the lemmas; culms forming corms.
→ 21
21. Panicle branches 3-11 cm long, divergent to reflexed, flexuous; lowest rachilla internodes 2-3 mm long
M. geyeri
21. Panicle branches 2-6.5 cm long, usually appressed to ascending, straight, sometimes strongly divergent and flexuous; lowest rachilla internodes 1-2 mm long.
→ 22
22. Ligules 0.1-2 mm long; glumes usually less than 1/2 the length of the spikelets; corms not attached directly to the rhizomes
M. spectabilis
22. Ligules 2-6 mm long; glumes from (1/2)2/3 as long as to equaling the spikelets; corms almost sessile, directly attached to the rhizomes
M. bulbosa
Source FNA vol. 24, p. 95. FNA vol. 24, p. 88. Author: Mary E. Barkworth;.
Parent taxa Poaceae > subfam. Pooideae > tribe Meliceae > Melica Poaceae > subfam. Pooideae > tribe Meliceae
Sibling taxa
M. altissima, M. aristata, M. bulbosa, M. californica, M. ciliata, M. frutescens, M. fugax, M. geyeri, M. harfordii, M. imperfecta, M. montezumae, M. mutica, M. nitens, M. porteri, M. spectabilis, M. stricta, M. subulata, M. torreyana
Subordinate taxa
M. altissima, M. aristata, M. bulbosa, M. californica, M. ciliata, M. frutescens, M. fugax, M. geyeri, M. harfordii, M. imperfecta, M. montezumae, M. mutica, M. nitens, M. porteri, M. smithii, M. spectabilis, M. stricta, M. subulata, M. torreyana
Name authority (Porter ex A. Gray) Vasey L.
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