Melica bulbosa |
Poaceae tribe Meliceae |
|||||||||||||
---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
bulbous oniongrass, onion grass, western onion grass |
|
|||||||||||||
Habit | Plants loosely cespitose, rhizomatous. | Plants usually perennial, sometimes annual; cespitose, sometimes rhizomatous. | ||||||||||||
Culms | 29-100 cm, forming corms, corms almost sessile on the connecting rhizomes; internodes scabridulous above the nodes. |
annual, not woody, not branching above the base; internodes hollow. |
||||||||||||
Sheaths | usually scabridulous, sometimes sparsely pilose; ligules 2-6 mm; blades 1.5-5 mm wide, abaxial surfaces scabridulous, adaxial surfaces with hairs. |
closed for their whole length or almost so; collars without tufts of hair on the sides; auricles sometimes present; ligules hyaline, glabrous, often lacerate, occasionally ciliate, those of the lower and upper cauline leaves usually similar; pseudopetioles absent; blades linear to narrowly lanceolate, venation parallel, cross venation sometimes evident; cross sections non-Kranz, without arm or fusoid cells; epidermes without microhairs, sometimes papillate. |
||||||||||||
Panicles | 7-30 cm; branches 2-6.5 cm, appressed, usually straight, with 1-5 spikelets; pedicels straight; disarticulation above the glumes. |
|||||||||||||
Inflorescences | terminal panicles or racemes; disarticulation above the glumes and beneath the florets or below the glumes. |
|||||||||||||
Spikelets | 6-24 mm, with 4-7 bisexual florets, base of the distal florets concealed at anthesis; rachilla internodes 1-2 mm, not swollen when fresh, not wrinkled when dry. |
2.5-60 mm, not viviparous, slightly to strongly laterally compressed, with 1-30 florets, proximal florets bisexual, distal 1-3 florets usually sterile, sometimes pistillate, sometimes reduced and amalgamated into a knob- or club-shaped rudiment; rachillas prolonged beyond the base of the distal floret. |
||||||||||||
Glumes | from (1/2)2/3 as long as to equaling the spikelets; lower glumes 5.5-10.5 mm long, 2-3 mm wide, 3-5-veined; upper glumes 6-14 mm long, 2.3-3.5 mm wide, 5-7-veined; lemmas 6-12 mm, glabrous, smooth or scabrous, 7-11-veined, veins prominent, apices emarginate to acute, unawned; paleas about 3/4 the length of the lemmas; anthers 3, 1.5-4 mm; rudiments 1.5-5 mm, truncate to tapering, sometimes resembling the bisexual florets in shape. |
exceeded by the distal florets, shorter than to longer than the adjacent lemmas, mostly membranous, scarious distally, 1-11-veined, apices usually rounded to acute; florets laterally or dorsally compressed; calluses blunt, glabrous or with hairs; lemmas of sexual florets rectangular or ovate, mostly membranous, scarious distally, often with a purplish band adjacent to the scarious apices, (4)5-15-veined, veins not converging distally, often prominent, unawned or awned, awns not branched, apices entire to bilobed or bifid, awns straight, subterminal or from the sinuses; paleas from shorter than to longer than the lemmas, similar in texture, 2-veined, veins keeled, sometimes winged; lodicules 2, fleshy, usually connate into a single structure, without a membranous wing, truncate, not ciliate, not or scarcely veined; anthers 1, 2, or 3; ovaries glabrous; styles 2-branched, bases persistent, branches plumose distally. |
||||||||||||
Caryopses | ovoid to ellipsoid, longitudinally grooved or not; hila usually linear; embryos less than 1/3 as long as the caryopses. |
|||||||||||||
x | = (8)9, 10. |
|||||||||||||
2n | =18. |
|||||||||||||
Melica bulbosa |
Poaceae tribe Meliceae |
|||||||||||||
Distribution |
CA; CO; ID; MT; NV; OR; SD; TX; UT; WA; WY; BC; SK
|
|||||||||||||
Discussion | Melica bulbosa grows from 1370-3400 m, mostly in open woods on dry, well-drained slopes and along streams. It is restricted to the western half of the Flora region. Two records from Texas, in Jeff Davis and Sutton counties, have not been verified. Melica bulbosa differs from M. spectabilis in its sessile corm and longer glumes. In addition, in M. bulbosa the spikelets have purplish bands which appear to be concentrated towards the apices; in M. spectabilis the bands appear more regularly spaced. It differs from M. californica in its more narrowly acute spikelets, more strongly colored lemmas, and lack of corms, and from M. fugax in not having swollen rachilla internodes. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
There are approximately 130 species and 8 or 9 genera in the Meliceae. Four of the genera are monotypic. Melica and Glyceria, the two largest genera, are well represented in North America. Pleuropogon and Schizachne are primarily North American, but extend into eastern Asia. Molecular studies (e.g., Soreng and Davis 2000; Grass Phylogeny Working Group 2001) show the tribe to be monophyletic and somewhat basal within the Pooideae. Members of the tribe are most easily recognized by the combination of closed leaf sheaths, scarious lemma apices, and non-converging lemma veins. The tribe also differs from other tribes in the Pooideae in having 2 unwinged lodicules that are usually connate into a single structure, and a base chromosome number of 9 or 10. Catabrosa and Briza, whose inclusion in the tribe was suggested by the preliminary results of Mejia-Saules and Bisby (2000), have more membranous lemma margins and free, winged lodicules. Briza also has open leaf sheaths and more convergent lemma veins. Their inclusion is not supported by the molecular data. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
||||||||||||
Key |
|
|||||||||||||
Source | FNA vol. 24, p. 91. | FNA vol. 24, p. 67. | ||||||||||||
Parent taxa | Poaceae > subfam. Pooideae > tribe Meliceae > Melica | Poaceae > subfam. Pooideae | ||||||||||||
Sibling taxa | ||||||||||||||
Subordinate taxa | ||||||||||||||
Synonyms | M. bulbosa var. intonsa, M. bulbosa var. inflata, Bromelica bulbosa | |||||||||||||
Name authority | Geyer ex Porter & J.M. Coult. | Endl. | ||||||||||||
Web links |
|