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climbing snapdragon, roving sailor, violet twining snapdragon

maurandella, snapdragon vine

Habit Perennials vines, 50–300 cm. Herbs, perennial; caudex woody.
Stems

climbing, glabrous.

Leaves

glabrous;

petiole 5–45 mm;

blade hastate to sagittate, 5–30 × 4–35 mm.

cauline, alternate;

petiole twining;

blade fleshy, not leathery, margins entire.

Inflorescences

axillary, flowers solitary;

bracts absent.

Pedicels

ascending, winged, 10–45 mm, glabrous.

present;

bracteoles absent.

Flowers

sepals subequal, adaxial lateral pair falcate, 9–14 × 2–3 mm, apex acute, surfaces glabrous;

corolla tube 13–17 mm, glabrous externally, throat closed, palate inflated and streaked with white, yellow, or violet, puberulent to villous, lobes rounded, 4–8 mm, abaxial lobes projecting, adaxial erect;

stamens included, filaments incurved, yellow-villous basally, abaxial 9–12 mm, adaxial 11–14 mm, pollen sacs oblong;

ovary septum V-shaped, locules unequal;

style terete, included, 10–13 mm, stigma recurved.

bisexual;

sepals 5, distinct, lanceolate, calyx bilaterally symmetric, campanulate;

corolla blue to violet, pink, or red, bilaterally symmetric, bilabiate and personate, tubular, tube base not spurred or gibbous, lobes 5, abaxial 3, adaxial 2;

stamens 4, basally adnate to corolla, didynamous, filaments glandular-hairy;

staminode 1, filamentous;

ovary 2-locular, placentation axile;

stigma 2-lobed.

Fruits

capsules, dehiscence loculicidal.

Capsules

globular, 7–10 mm, glabrous, larger locule dehiscing first.

Seeds

1–1.5 mm, surfaces cristate-tuberculate.

80–150, dark brown to black, ovoid to oblong-polygonal, wings absent.

x

= 12.

2n

= 24.

Maurandella antirrhiniflora

Maurandella

Phenology Flowering Feb–Dec.
Habitat Canyons, rocky slopes, arroyos, coastal dunes, desert flats on siliceous, calcareous, gypseous substrates.
Elevation 0–1800(–2600) m. (0–5900(–8500) ft.)
Distribution
from FNA
AZ; CA; NM; NV; TX; Mexico
[WildflowerSearch map]
[BONAP county map]
from USDA
sw United States; sc United States; Mexico
[BONAP county map]
Discussion

Maurandella antirrhiniflora has been treated either as one of two species in Maurandella (along with M. hederifolia) (W. Rothmaler 1943; D. A. Sutton 1988), or as one species with M. hederifolia treated at subspecific rank (W. J. Elisens 1985). Maurandella antirrhiniflora is recognized here without infraspecific variants to be consistent with the pattern of interspecific variation in Antirrhineae. Variable floral coloration is present in central Arizona and California, with pink and red corolla morphs occurring in discrete populations from the widespread blue to violet morphs.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 2 (1 in the flora).

Maurandella most commonly has been recognized as a genus segregated from Maurandya Ortega or as a section within Maurandya, based on distinctive characteristics: personate corolla, globular capsule, V-shaped septum, and marked asymmetry of the locules. The V-shaped septum results in unequal locule volumes, which is unique in Antirrhineae. Based on this unique combination of characteristics, Maurandella is recognized here as a genus distinct from Epixiphium and Maurandya.

Initial molecular phylogenetic studies indicated a sister group relationship between species of Maurandella and Maurandya (M. Ghebrehiwet et al. 2000); they lacked the taxon sampling needed to resolve relationships among closely related species in Epixiphium, Maurandella, and Maurandya (C. E. Freeman and R. Scogin 1999; P. Vargas et al. 2004). The more complete ITS sampling by M. Fernández-Mazuecos et al. (2013) placed both genera in the Cymbalaria clade, but not as sister taxa; Maurandya was sister to Lophospermum, whereas Maurandella was nested in a more basal position within a seven-taxon subclade comprising genera in subtribe Maurandyinae.

Maurandella hederifolia Rothmaler is known from northern Mexico.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 35. FNA vol. 17, p. 35. Author: Wayne J. Elisens.
Parent taxa Plantaginaceae > Maurandella Plantaginaceae
Subordinate taxa
M. antirrhiniflora
Synonyms Maurandya antirrhiniflora, Antirrhinum antirrhiniflora, Asarina antirrhiniflora Antirrhinum
Name authority (Humboldt & Bonpland ex Willdenow) Rothmaler: Feddes Repert. Spec. Nov. Regni Veg. 52: 27. (1943) (A. Gray) Rothmaler: Feddes Repert. Spec. Nov. Regni Veg. 52: 26. (1943)
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