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bitter cassava, cassava, manioc, tapioca

palo mulato, spike manihot

Habit Shrubs, 1–4 m. Herbs or subshrubs, to 1 m.
Roots

thickened.

thickened.

Stems

erect, terete when young;

nodes conspicuously swollen;

leaf and stipule scars elevated, especially on older stems.

lax (often leaning on other vegetation), terete when young;

nodes not swollen;

leaf and stipule scars not elevated.

Leaves

persistent;

stipules lanceolate, entire;

petiole 3–20 cm;

blade basally attached, usually 3–10-lobed, sometimes unlobed, lobes without secondary lobes, median lobe 5–18 cm, margins neither thickened nor revolute, entire to ± repand, apex acuminate, surfaces glabrous or hairy, abaxial finely reticulate.

stipules lanceolate, entire;

petiole 2–10 cm;

blade basally attached to subpeltate, 5-lobed, lobes with acute secondary lobes near base, median lobe 2–10 cm, margins thickened and revolute, remotely serrate, apex acute to acuminate (bristle-tipped), surfaces glabrous, abaxial smooth.

Inflorescences

axillary, panicles, 2–10 cm.

terminal, racemes, 25 cm.

Pedicels

staminate 2–4 mm; pistillate 20 mm in fruit, straight.

staminate 1–5 mm; pistillate 10–20 mm in fruit, downcurved.

Staminate flowers

calyx campanulate, 10–15 mm, lobes erect or spreading;

stamens 10.

calyx campanulate to conic, 8–13 mm, lobes reflexed;

stamens 10.

Capsules

1.5 cm, usually winged.

1.5 cm, smooth, not winged.

Seeds

subglobose to oblong, 12 mm.

oblong, 10 mm.

2n

= 36.

Manihot esculenta

Manihot subspicata

Phenology Flowering year-round, mostly fall and winter. Flowering Jun–Aug.
Habitat Disturbed areas, spreading from cultivation. Savannahs and grasslands with scattered shrubs and trees.
Elevation 0–200 m. (0–700 ft.) 30–60 m. (100–200 ft.)
Distribution
from FNA
AL; FL; TX; South America (Brazil) [Introduced in North America; introduced widely in tropical and subtropical regions worldwide]
[WildflowerSearch map]
[BONAP county map]
from FNA
TX; Mexico (Coahuila, Nuevo León, Tamaulipas)
[BONAP county map]
Discussion

The enlarged storage roots of Manihot esculenta yield a starchy staple, now much consumed in tropical regions around the world. Tapioca, a pelletized and partially hydrolyzed form of cassava starch, is the chief form of consumption in temperate regions. Multiple cultivars are known. These are generally characterized as bitter (containing cyanogenic glycosides, which must be removed before consumption) or sweet (cyanogenic glycosides absent or at low levels). A form with variegated leaves is sometimes grown for ornament. Cassava was cultivated throughout the Neotropics in pre-Columbian times. As a root crop with poor storage qualities adapted to humid regions, archeological remains are few, leading to much speculation in the literature about the origin of this important crop. Molecular data reported by K. Olsen and B. A. Schaal (1999, 2001), indicate that cultivated cassava constitutes M. esculenta subsp. esculenta, derived by artificial selection from its sole wild ancestor, M. esculenta subsp. flabellifolia (Pohl) Ciferri from the southern border of the Amazon basin. Under this classification, all North American plants belong to subsp. esculenta.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Although D. J. Rogers and S. G. Appan (1973) characterized leaf blades of Manihot subspicata as peltate, seldom are leaves unambiguously so. Typically, just the thickened margins of lateral lobes are confluent across the distal end of the petiole. Though relatively common in northern Mexico, M. subspicata is known in Texas only from the vicinity of Lake Corpus Christi (Jim Wells and Live Oak counties); whether native or introduced there is unresolved. In Mexico, M. subspicata appears to be tolerant of disturbance, frequently colonizing roadsides and similar habitats.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 194. FNA vol. 12, p. 196.
Parent taxa Euphorbiaceae > Manihot Euphorbiaceae > Manihot
Sibling taxa
M. angustiloba, M. davisiae, M. grahamii, M. subspicata, M. walkerae
M. angustiloba, M. davisiae, M. esculenta, M. grahamii, M. walkerae
Name authority Crantz: Inst. Rei Herb. 1: 167. (1766) D. J. Rogers & Appan: in Organization for Flora Neotropica, Fl. Neotrop. 13: 62, figs. 19D, 20A–C. (1973)
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