Manfreda virginica |
Agavaceae |
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false aloe, rattlesnake master, Virginian agave |
agave or century plant family, century plant family |
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Habit | Plants usually perennial, occasionally epiphytic, sometimes monocarpic or polycarpic, monoecious, dioecious, or polygamodioecious, small to gigantic, sometimes arborescent, usually scapose. | |||||||||||||||||||||||||||||||||
Rhizomes | cylindrical. |
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Stems | subterranean or aboveground, sometimes branched. |
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Leaves | spreading, semisucculent, 8–40(–47) × 0.5–6.5(–9.3) cm; blade usually spotted or speckled with maroon, shallowly channeled, oblanceolate to linear-lanceolate, margins entire or with cartilaginous prickles. |
simple, annual or long-lived, in terminal rosettes or occasionally cauline, sessile or occasionally pseudo-petiolate; blade linear, lanceolate, oblanceolate, ovate, or elliptic, fibrous, thin and flexible, thick and rigid or succulent, or fibrous, often glaucous, margins entire, serrulate, dentate, denticulate, corneous, or filiferous, apex rigid or flexible, sometimes pungent, often with short or long spine. |
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Scape | 4.5 13.8 dm. |
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Inflorescences | 14–68 cm, bearing 10–61 closely spaced flowers. |
terminal or axillary spikes, racemose or paniculate, sometimes umbellate, bracteate, often huge; bracts ascending or erect, occasionally reflexed, leaflike proximally, scalelike distally. |
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Flowers | sessile or pedicellate, nearly erect, slender, with sweet, fruity odor; tepals green; perianth tube 0.9–2.3 × 0.3–0.6 cm; limb lobes erect, 0.4–0.8 cm; filaments inserted near base of tube, bent in bud, exceeding tube by 1.2–3.1 cm; ovary 4–10 mm; style shorter than stamens, exceeding tube by 0.6–2.3 cm; stigma white, 3-lobed, lobes reflexed. |
6-merous, bisexual or functionally unisexual; perianth of 2 similar petallike whorls, semisucculent; tepals distinct or connate into tube, apex glandular or glandular-pubescent; stamens included or exserted; filaments often broadened and succulent, glabrous, pubescent, or papillose; anthers versatile, dehiscence longitudinal; ovary superior or inferior, 3-locular or occasionally 1-locular, 3-angled, ovoid, or cylindrical, with axillary or rarely parietal placentation; style included or exserted; stigmas 1 or 3, 3-lobed or capitate; pedicel usually distinct, articulate or not, rarely absent. |
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Fruits | occasionally baccate, usually capsular and sometimes winged or lobed, or indehiscent and dry or fleshy. |
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Capsules | globose, 1–1.7 cm diam. 2n = 60. |
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Seeds | 1–3(–many) per locule, flattened, 3-angled, hemispheric, ovoid, obovoid, or globose. |
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Manfreda virginica |
Agavaceae |
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Phenology | Flowering summer–late summer, rarely in spring; fruiting late summer–early fall. | |||||||||||||||||||||||||||||||||
Habitat | Glades and open woods, on rocky and sandy soils, often on slopes | |||||||||||||||||||||||||||||||||
Elevation | 0–600 m (0–2000 ft) | |||||||||||||||||||||||||||||||||
Distribution |
AL; AR; FL; GA; IL; IN; KY; LA; MO; MS; NC; OH; OK; SC; TN; TX; VA; WV; Mexico (Nuevo León, Tamaulipas)
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Worldwide; primarily arid; semitropical; subtropical; and tropical regions |
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Discussion | Leaf shape and size in Manfreda virginica vary with soil type, amount of shade, length of cold period, and position of leaf in the rosette. Speckles and spots occur frequently on some leaves in most populations, and some authors have used the informal designation “forma tigrina for such variants. Pollination is primarily by sphinx moths (S. E. Verhoek 1978). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera 17 or 18, species ca. 550 (9 genera, 84 species in the flora; 2 genera, 6 species introduced). There is little agreement on the treatment of Agavaceae. The group containing Agave, Yucca, Furcraea, Hesperaloe, and Manfreda generally has been accepted as the core of Agavaceae, or as subfamilies Agavoideae and Yuccoideae, but treatment of Dracaena, Sansevieria, Cordyline, Nolina, and Dasylirion has been varied. A. L. Takhtajan (1987) and R. F. Thorne (1992b) placed these genera in Dracaenaceae but treated them at different levels. Takhtajan located them in the subfamily Dracaenoideae within sections Nolineae (Nolina and Dasylirion), Sansevierieae (Sansevieria), and Dracaeneae (Cordyline and Dracaena). Thorne, on the other hand, placed these same groupings at the subfamily level. R. M. T. Dahlgren et al. (1985) recognized them as separate families, Nolinaceae (Nolina and Dasylirion), Dracaenaceae (Sansevieria and Dracaena), and Asteliaceae (Cordyline), in addition to the Agavaceae (Yucca, Hesperaloe, Agave, Manfreda, and Furcraea). A. Cronquist (1981) based his broadly circumscribed Agavaceae on a common xerophytic habit. However, the karyotype of 5 long and 25 short chromosomes for the Agavoideae and Yuccoideae is distinct from the karyotypes of the other subfamilies that Cronquist included in the Agavaceae. Current research on the phylogenetics of moncotyledons, using DNA sequences of rbcL, support the separation of Dracaena, Nolina, and Dasylirion from Agavaceae (M. R. Duvall et al. 1993b). We believe that a broad interpretation of the Agavaceae unites groups that should be recognized as separate. Many genera in Agavaceae are economically important. All genera in the Agavoideae and Yuccoideae contain steroidal sapogenins; some have been used in folk medicine, and locally and commercially as soap (G. Blunden et al. 1978; S. E. Verhoek 1978; M. Wall et al. 1957). They provide fibers for cordage, baskets, and hats, as well as food and drink for many indigenous peoples of the southwestern United States (H. S. Gentry 1982). They are also used as commercial fiber and beverage crops in Latin America and the Old World (H. Brucher 1989). In the southern United States, some species in each genus are cultivated and represented in the flora, and at least one species of Yucca is now grown as far north as Canada. Collectors should record the uses of these plants in their notes along with the critical information on plant habit and morphology. Photographs are often important tools for the identification of these plants, and, with the advent of digital cameras, are now much easier to obtain and process. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 26, p. 463. | FNA vol. 26, p. 413. | ||||||||||||||||||||||||||||||||
Parent taxa | Agavaceae > Manfreda | |||||||||||||||||||||||||||||||||
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Synonyms | Agave virginica, Agave lata, Agave tigrina, Agave virginica var. tigrina, M. tigrina, M. virginica subsp. lata, M. virginica var. tigrina, Polianthes lata, Polianthes virginica | |||||||||||||||||||||||||||||||||
Name authority | (Linnaeus) Salisbury ex Rose: Contr. U.S. Natl. Herb. 8: 19. (1903) | Dumortier | ||||||||||||||||||||||||||||||||
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