Mammillaria tetrancistra |
Mammillaria mainiae |
|
|---|---|---|
| common fish hook cactus |
counterclockwise fishhook cactus, counterclockwise nipple cactus |
|
| Habit | Plants branched; branches 1–several. | Plants branched or unbranched; branches 0–several, not numerous, seldom rooting. |
| Roots | fleshy taproots, to 24 cm, 5–8 cm diam. |
diffuse, upper portion not enlarged. |
| Stems | cylindric to ovoid-cylindric, commonly 5–15(–25) × 3.5–7(–10) cm, flaccid; tubercles 4 mm diam.; axils short woolly; cortex and pith mucilaginous; latex absent. |
nearly spheric, 2.5–12 × 3.5–7 cm, firm; tubercles 9–10.5(–18) × 3–9 mm; axils appearing naked; cortex and pith not mucilaginous; latex absent. |
| Spines | 21–64 per areole, dark or light colored, depending largely on substrate color, glabrous (to hoary); radial spines 30–46(–60) per areole, white, bristlelike, 6–10 × 0.09–0.15 mm, stiff; central spines 1–3(–4) per areole, porrect or strongly projecting, usually hooked, (6–)13–18(–25) × (0.2–)0.3(–0.4) mm; subcentral spines several, often 12+ per areole, radiating in all directions, often resembling supplementary ring of radial spines, barely distinguishable from radial spines, stouter, longer and dark tipped or purplish. |
ca. 9–16 per areole, brightly colored than M. grahamii, yellowish, pale pinkish tan, or brown (smaller spines paler), tipped dark chestnut brown to blackish, glabrous, sometimes ± pubescent when young; radial spines (8–)10–15 per areole, bristlelike, 6–10(–12) × 0.13–0.23 mm, stiff; central spines 1(–2) per areole, porrect, hooked, 11–20 × 0.2–0.4 mm; subcentral spines 0(–2) per areole, adaxial to central spines. |
| Flowers | 2.5 × 2.5–3.5 cm; outermost tepal margins long fringed; inner tepals pink to rose-purple, margins sometimes paler or white, at least proximally, 24–26 × 4 mm; stigma lobes yellow-green to green. |
(1–)2–3 × 1.2–2 cm; outermost tepal margins densely fringed, fringes 0.4 mm; inner tepals pinkish white with sharply defined magenta midstripes; stigma lobes bright red to red-purple, orange, or magenta, (3–)7–9 mm. |
| Fruits | bright red, ellipsoid or cylindric to clavate, (8–)15–30 × 5–10 mm, juicy only in fruit walls; floral remnant quickly deciduous, leaving conspicuous abscission scar. |
bright orange-red, spheric to obovoid, 5–7 × 4–4.5 mm, level with or beneath spines, juicy only in fruit walls; floral remnant weakly persistent. |
| Seeds | black, conspicuously strophiolate, 1.4–2.4 × 1.4 mm, pitted and rugose; testa hard; anticlinal cell walls straight (not undulate); interstices narrower than pit diameters; pits bowl-shaped; strophiole tan, large, corky. |
black, 1–1.2 × 0.8–1 mm, pitted; testa hard; anticlinal cell walls straight; interstices conspicuously wider than pit diameters; pits bowl-shaped. |
| 2n | = 22. |
= 22. |
Mammillaria tetrancistra |
Mammillaria mainiae |
|
| Phenology | Flowering Apr, Jul; fruiting Feb–Apr, Sep–Oct. | Flowering summer. |
| Habitat | Mojave and Sonoran Deserts, alluvium and outcrops, valley floors, hills, mountainsides | Sonoran desert, grasslands, bajadas, valleys, washes, alluvial fans [subtropical woodlands, tropical deciduous forests] |
| Elevation | 100-1500 m [300-4900 ft] | [200-]600-1200 m [[700-]2000-3900 ft] |
| Distribution |
AZ; CA; NV; UT; Mexico (Baja California, Sonora) |
AZ; Mexico (Sinaloa, Sonora) |
| Discussion | Mammillaria tetrancistra extends farther into hyper-arid California deserts than any other species of Mammillaria. Without the unique seeds, its identification requires detailed comparison with both Mammillaria grahamii and M. viridiflora. Although M. viridiflora is ecogeographically segregated (more mesophytic), the other taxa grow intermingled at many sites in southwestern Arizona. Pushing the side of the stems with a stick or stone allows crude field identification for two commonly confused species: stems of M. tetrancistra are soft and flabby, whereas stems of M. grahamii are firm. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The tendency for all spine hooks on plants to be oriented in same direction is not unique to Mammillaria mainiae. This uncommon and poorly known species is restricted in the flora area to the relatively mesic eastern edge of the Sonoran Desert, in western bajadas of the Baboquivari Mountains, Arizona. Mammillaria mainiae is not known from Nogales, Arizona, contrary to L. D. Benson (1969, 1982); it was originally discovered in Mexico somewhere south of Nogales. Mammillaria wrightii var. wilcoxii, which grows all around Nogales, Arizona, is easily misidentified as M. mainiae. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
| Parent taxa | ||
| Sibling taxa | ||
| Name authority | Engelmann: Amer. J. Sci. Arts, ser. 2, 14: 337. (1852) | K. Brandegee: Zoë 5: 31. (1900) |
| Source | FNA vol. 4, p. 251. | FNA vol. 4, p. 253. |
| Web links | ||