Mammillaria tetrancistra |
Mammillaria macdougalii |
|
|---|---|---|
| common fish hook cactus |
cream cactus |
|
| Habit | Plants branched; branches 1–several. | Plants unbranched, deeply seated in substrate. |
| Roots | fleshy taproots, to 24 cm, 5–8 cm diam. |
short, obconic taproots; secondary roots diffuse. |
| Stems | cylindric to ovoid-cylindric, commonly 5–15(–25) × 3.5–7(–10) cm, flaccid; tubercles 4 mm diam.; axils short woolly; cortex and pith mucilaginous; latex absent. |
usually flat-topped (in old age or under dense brush hemispheric), 4–20 × 9-25(–30) cm, firm; tubercles 10–17 × 6–8.5 mm; axils woolly, wool 5–8 mm, bristles absent; cortex and pith not mucilaginous; latex sticky white, abundant in healthy tissue throughout cortex of stem, tubercles, and sometimes flower receptacles. |
| Spines | 21–64 per areole, dark or light colored, depending largely on substrate color, glabrous (to hoary); radial spines 30–46(–60) per areole, white, bristlelike, 6–10 × 0.09–0.15 mm, stiff; central spines 1–3(–4) per areole, porrect or strongly projecting, usually hooked, (6–)13–18(–25) × (0.2–)0.3(–0.4) mm; subcentral spines several, often 12+ per areole, radiating in all directions, often resembling supplementary ring of radial spines, barely distinguishable from radial spines, stouter, longer and dark tipped or purplish. |
11–14(–16) per areole, white to straw yellow, glabrous; radial spines 10–12 per areole, whitish to pale yellowish tan, needlelike, 20 mm, stiff; central spines 1–2(–4) per areole, ± porrect or 1 ascending and 1 descending, straight or slightly curved, 10 mm; subcentral spines 0. |
| Flowers | 2.5 × 2.5–3.5 cm; outermost tepal margins long fringed; inner tepals pink to rose-purple, margins sometimes paler or white, at least proximally, 24–26 × 4 mm; stigma lobes yellow-green to green. |
3–4 × 2.5–3.5 cm; outermost tepal margins densely minutely fimbriate (at 10x); inner tepals greenish white to greenish yellow, 14–17(–25) × 2–3 mm; stigma lobes yellow-green to pale green, 2–3 mm. |
| Fruits | bright red, ellipsoid or cylindric to clavate, (8–)15–30 × 5–10 mm, juicy only in fruit walls; floral remnant quickly deciduous, leaving conspicuous abscission scar. |
whitish green proximally, pale green distally with pale rose-purple on sunlit portions, obovoid to clavate, 15–26 × 7.5–15 mm, juicy only in fruit walls; floral remnant persistent. |
| Seeds | black, conspicuously strophiolate, 1.4–2.4 × 1.4 mm, pitted and rugose; testa hard; anticlinal cell walls straight (not undulate); interstices narrower than pit diameters; pits bowl-shaped; strophiole tan, large, corky. |
reddish brown or orange, 1.1–1.3 × 0.9 × 0.8 mm, pitted; testa intermediate, anticlinal cell walls undulate, interstices much narrower than pit diam., pits deeply concave, elongate. |
| 2n | = 22. |
= 22. |
Mammillaria tetrancistra |
Mammillaria macdougalii |
|
| Phenology | Flowering Apr, Jul; fruiting Feb–Apr, Sep–Oct. | Flowering Mar–May; fruiting Oct–Mar. |
| Habitat | Mojave and Sonoran Deserts, alluvium and outcrops, valley floors, hills, mountainsides | Rocky slopes and ridges, ecotone between Sonoran desert scrub and chaparral, oak woodlands, grasslands |
| Elevation | 100-1500 m [300-4900 ft] | 1100-1800 m [3600-5900 ft] |
| Distribution |
AZ; CA; NV; UT; Mexico (Baja California, Sonora) |
AZ; Mexico (Sonora) |
| Discussion | Mammillaria tetrancistra extends farther into hyper-arid California deserts than any other species of Mammillaria. Without the unique seeds, its identification requires detailed comparison with both Mammillaria grahamii and M. viridiflora. Although M. viridiflora is ecogeographically segregated (more mesophytic), the other taxa grow intermingled at many sites in southwestern Arizona. Pushing the side of the stems with a stick or stone allows crude field identification for two commonly confused species: stems of M. tetrancistra are soft and flabby, whereas stems of M. grahamii are firm. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Mammillaria macdougalii is the characteristic flat-topped mammillaria of the mountains near Tucson and Nogales, Arizona. Records from farther east are all (insofar as can be determined) misidentifications of M. heyderi var. bullingtoniana. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
| Parent taxa | ||
| Sibling taxa | ||
| Synonyms | M. gummifera var. macdougalii, M. heyderi var. macdougalii | |
| Name authority | Engelmann: Amer. J. Sci. Arts, ser. 2, 14: 337. (1852) | Rose: in L. H. Bailey, Stand. Cycl. Hort. 4: 1982. (1916) |
| Source | FNA vol. 4, p. 251. | FNA vol. 4, p. 257. |
| Web links | ||