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biznaga China, Heyder's pincushion cactus, little nipple cactus

biznaga de Wright, Wright's nipple cactus, Wright's pincushion cactus

Habit Plants unbranched, protruding relatively little above soil. Plants usually unbranched.
Roots

obconic taproots;

secondary roots diffuse.

upper portion of primary root somewhat thickened and succulent proximally, otherwise fibrous.

Stems

top-shaped, flat-topped (aerial part sometimes hemispheric in old age or in dense subtropical vegetation), protruding above ground 0–2 × (4–)7.5–15 cm, firm;

tubercles 9–15(–20) × 3–7 mm;

axils with short wool, bristles absent;

cortex and pith not mucilaginous;

latex abundant in healthy tissue throughout cortex of stem, tubercles, and sometimes flower receptacle, sticky, white.

flat-topped or spheric, 4–8 × 4–8 cm, ± flaccid;

tubercles 6–24 mm;

axils appearing naked;

cortex and pith mucilaginous;

latex absent.

Spines

(8–)10–18(–27) per areole, usually brownish, darker at tip, glabrous;

radial spines (8–)10–22(–26) per areole, white to white-and-brown or brown, needlelike, 6–15(–16) mm, stiff, abaxial spines longest;

central spines (0–)1(–4) per areole, porrect or ascending, not hooked, (0.5–)2–8 × 0.15–0.45 mm;

subcentral spines 0.

9–31(–34) per areole, white, usually tipped brown;

radial spines 8–30 per areole, in 1 series, glabrous (puberulent in first year flowering plants);

lateral spines bristlelike, 7–11 mm, stiff, longest and thickest of spines;

adaxial spines often darker;

central spines 1–4(–7) per areole, porrect or strongly projecting, 1 or all hooked, (5–)12–14(–21) mm;

subcentral spines 0.

Flowers

1.9–3.8 × 1.5–3 cm;

outermost tepal margins entire;

inner tepals white, greenish or cream to pale pink, with tan, pink, greenish, or brownish midstripes, 11–19 × 2–2.5 mm;

stigma lobes externally green, internally green or red (or pink), 2.5–3 mm.

2.5–3.5(–5.2) × 2.2–4.5(–7.5) cm;

outer tepal margins conspicuously fringed;

inner tepals rose-pink or magenta [white], margins often paler;

stigma lobes yellow or pale green (rarely reddish).

Fruits

brilliant red: scarlet, carmine, or crimson, obovoid to clavate, 10–35(–40) × 5–8 mm, juicy only in fruit walls;

floral remnant weakly persistent.

green or dull purple, spheric to ovoid or obovoid, (9–)13–20(–28) × (6–)7–1.9(–26) mm, juicy throughout;

floral remnant persistent.

Seeds

reddish brown, sometimes yellowish when fresh, 1–1.2 mm, deeply pitted;

testa thin, relatively flexible;

anticlinal cell walls sinuate, interstices narrower than pit diameters;

pits cavernous or deeply concave.

black, 1.3–1.5 mm, pitted;

testa hard, anticlinal cell walls straight (not undulate);

interstices conspicuously narrower than pit diameters;

pits bowl-shaped.

2n

= 22.

= 22.

Mammillaria heyderi

Mammillaria wrightii

Distribution
from FNA
AZ; NM; OK; TX; n Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM; TX; Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Green fruits of Mammillaria heyderi with fully mature, viable seeds precede the ripe (elongate) fruits by six months to a year.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 3 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Central spines 0.15-0.35 mm diam.; radial spines (7-)13-17(-26) per areole, abaxial radial spines 6-11(-16) mm; e of El Paso, Texas, New Mexico, Oklahoma.
var. heyderi
1. Central spines 0.35-0.45 mm diam.; radial spines 10-14 per areole, abaxial radial spines 9-15 mm; El Paso, Texas, w to Arizona
var. bullingtoniana
1. Radial spines usually 8-15 per areole; fruits 12.5-26 mm diam
var. wrightii
1. Radial spines usually 16-30 per areole; fruits 6-15 mm diam
var. wilcoxii
Source FNA vol. 4, p. 255. FNA vol. 4, p. 250.
Parent taxa Cactaceae > subfam. Cactoideae > Mammillaria Cactaceae > subfam. Cactoideae > Mammillaria
Sibling taxa
M. dioica, M. grahamii, M. lasiacantha, M. macdougalii, M. mainiae, M. meiacantha, M. pottsii, M. prolifera, M. sphaerica, M. tetrancistra, M. thornberi, M. viridiflora, M. wrightii
M. dioica, M. grahamii, M. heyderi, M. lasiacantha, M. macdougalii, M. mainiae, M. meiacantha, M. pottsii, M. prolifera, M. sphaerica, M. tetrancistra, M. thornberi, M. viridiflora
Subordinate taxa
M. heyderi var. bullingtoniana, M. heyderi var. heyderi
M. wrightii var. wilcoxii, M. wrightii var. wrightii
Name authority Muehlenpfordt: Allg. Gartenzeitung 16: 20. (1848) Engelmann: Proc. Amer. Acad. Arts 3: 262. (1856)
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