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biznaga China, Heyder's pincushion cactus, little nipple cactus

green flower nipple cactus

Habit Plants unbranched, protruding relatively little above soil. Plants usually unbranched.
Roots

obconic taproots;

secondary roots diffuse.

upper portion of primary root somewhat thickened and succulent proximally, otherwise fibrous.

Stems

top-shaped, flat-topped (aerial part sometimes hemispheric in old age or in dense subtropical vegetation), protruding above ground 0–2 × (4–)7.5–15 cm, firm;

tubercles 9–15(–20) × 3–7 mm;

axils with short wool, bristles absent;

cortex and pith not mucilaginous;

latex abundant in healthy tissue throughout cortex of stem, tubercles, and sometimes flower receptacle, sticky, white.

flat-topped or spheric to short cylindric, ± flaccid;

tubercles (5–)6–15(–17) mm;

axils appearing naked;

cortex and pith mucilaginous;

latex absent.

Spines

(8–)10–18(–27) per areole, usually brownish, darker at tip, glabrous;

radial spines (8–)10–22(–26) per areole, white to white-and-brown or brown, needlelike, 6–15(–16) mm, stiff, abaxial spines longest;

central spines (0–)1(–4) per areole, porrect or ascending, not hooked, (0.5–)2–8 × 0.15–0.45 mm;

subcentral spines 0.

19–31(–34) per areole, usually white or brown-and-white (rarely bright reddish brown), glabrous (to hoary);

radial spines (13–)15–23(–31) per areole, in 1 series, bristlelike, stiff;

central spines 1–2(–4) per areole, porrect or strongly projecting, all hooked, (7.5–)10–20(–31) × 0.2–0.4 mm;

subcentral spines 0 [several in M. barbata].

Flowers

1.9–3.8 × 1.5–3 cm;

outermost tepal margins entire;

inner tepals white, greenish or cream to pale pink, with tan, pink, greenish, or brownish midstripes, 11–19 × 2–2.5 mm;

stigma lobes externally green, internally green or red (or pink), 2.5–3 mm.

(1.5–)2–3.5(–3.9) × (1.2–)1.8–3(–4.2) cm;

outermost tepal margins long fringed, reaching 1 mm;

inner tepals usually white, cream, pale tan, greenish white, or pale rose-pink, sometimes appearing pale orange proximally due to reflection of yellow anthers, usually with ± sharply defined pink midstripes (often suffused with brown, yellow, pink, red, purple, or green, darkest and brightest distally on most plants), (7.5–)12–17(–21) mm;

stigma lobes green or yellow-green.

Fruits

brilliant red: scarlet, carmine, or crimson, obovoid to clavate, 10–35(–40) × 5–8 mm, juicy only in fruit walls;

floral remnant weakly persistent.

green or purple, ovoid or obovoid, (6–)10–16(–22) × (4–)6–12(–13) mm, juicy throughout;

floral remnant persistent.

Seeds

reddish brown, sometimes yellowish when fresh, 1–1.2 mm, deeply pitted;

testa thin, relatively flexible;

anticlinal cell walls sinuate, interstices narrower than pit diameters;

pits cavernous or deeply concave.

chocolate brown with darker reticulation, 1.3 × 0.9 mm, pitted;

testa hard;

anticlinal cell walls straight;

interstices conspicuously narrower than pit diameters;

pits bowl-shaped.

2n

= 22.

= 22.

Mammillaria heyderi

Mammillaria viridiflora

Phenology Flowering spring; fruiting fall.
Habitat Semidesert grasslands, interior chaparral, pinyon-juniper and oak woodlands, crevices, boulders, canyon sides and gravelly igneous substrates
Elevation (800-)1400-2000 m ((2600-)4600-6600 ft)
Distribution
from FNA
AZ; NM; OK; TX; n Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM
Discussion

Varieties 2 (2 in the flora).

Green fruits of Mammillaria heyderi with fully mature, viable seeds precede the ripe (elongate) fruits by six months to a year.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Mammillaria viridiflora is closely related to M. barbata Engelmann of Mexico, which has priority if the species are combined.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Central spines 0.15-0.35 mm diam.; radial spines (7-)13-17(-26) per areole, abaxial radial spines 6-11(-16) mm; e of El Paso, Texas, New Mexico, Oklahoma.
var. heyderi
1. Central spines 0.35-0.45 mm diam.; radial spines 10-14 per areole, abaxial radial spines 9-15 mm; El Paso, Texas, w to Arizona
var. bullingtoniana
Source FNA vol. 4, p. 255. FNA vol. 4, p. 251.
Parent taxa Cactaceae > subfam. Cactoideae > Mammillaria Cactaceae > subfam. Cactoideae > Mammillaria
Sibling taxa
M. dioica, M. grahamii, M. lasiacantha, M. macdougalii, M. mainiae, M. meiacantha, M. pottsii, M. prolifera, M. sphaerica, M. tetrancistra, M. thornberi, M. viridiflora, M. wrightii
M. dioica, M. grahamii, M. heyderi, M. lasiacantha, M. macdougalii, M. mainiae, M. meiacantha, M. pottsii, M. prolifera, M. sphaerica, M. tetrancistra, M. thornberi, M. wrightii
Subordinate taxa
M. heyderi var. bullingtoniana, M. heyderi var. heyderi
Synonyms Neomammillaria viridiflora, M. orestera
Name authority Muehlenpfordt: Allg. Gartenzeitung 16: 20. (1848) (Britton & Rose) Boedeker: Mamm.-vergl.-Schlüssel, 36. (1933)
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