Mammillaria heyderi |
Mammillaria viridiflora |
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biznaga China, Heyder's pincushion cactus, little nipple cactus |
green flower nipple cactus |
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Habit | Plants unbranched, protruding relatively little above soil. | Plants usually unbranched. | ||||
Roots | obconic taproots; secondary roots diffuse. |
upper portion of primary root somewhat thickened and succulent proximally, otherwise fibrous. |
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Stems | top-shaped, flat-topped (aerial part sometimes hemispheric in old age or in dense subtropical vegetation), protruding above ground 0–2 × (4–)7.5–15 cm, firm; tubercles 9–15(–20) × 3–7 mm; axils with short wool, bristles absent; cortex and pith not mucilaginous; latex abundant in healthy tissue throughout cortex of stem, tubercles, and sometimes flower receptacle, sticky, white. |
flat-topped or spheric to short cylindric, ± flaccid; tubercles (5–)6–15(–17) mm; axils appearing naked; cortex and pith mucilaginous; latex absent. |
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Spines | (8–)10–18(–27) per areole, usually brownish, darker at tip, glabrous; radial spines (8–)10–22(–26) per areole, white to white-and-brown or brown, needlelike, 6–15(–16) mm, stiff, abaxial spines longest; central spines (0–)1(–4) per areole, porrect or ascending, not hooked, (0.5–)2–8 × 0.15–0.45 mm; subcentral spines 0. |
19–31(–34) per areole, usually white or brown-and-white (rarely bright reddish brown), glabrous (to hoary); radial spines (13–)15–23(–31) per areole, in 1 series, bristlelike, stiff; central spines 1–2(–4) per areole, porrect or strongly projecting, all hooked, (7.5–)10–20(–31) × 0.2–0.4 mm; subcentral spines 0 [several in M. barbata]. |
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Flowers | 1.9–3.8 × 1.5–3 cm; outermost tepal margins entire; inner tepals white, greenish or cream to pale pink, with tan, pink, greenish, or brownish midstripes, 11–19 × 2–2.5 mm; stigma lobes externally green, internally green or red (or pink), 2.5–3 mm. |
(1.5–)2–3.5(–3.9) × (1.2–)1.8–3(–4.2) cm; outermost tepal margins long fringed, reaching 1 mm; inner tepals usually white, cream, pale tan, greenish white, or pale rose-pink, sometimes appearing pale orange proximally due to reflection of yellow anthers, usually with ± sharply defined pink midstripes (often suffused with brown, yellow, pink, red, purple, or green, darkest and brightest distally on most plants), (7.5–)12–17(–21) mm; stigma lobes green or yellow-green. |
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Fruits | brilliant red: scarlet, carmine, or crimson, obovoid to clavate, 10–35(–40) × 5–8 mm, juicy only in fruit walls; floral remnant weakly persistent. |
green or purple, ovoid or obovoid, (6–)10–16(–22) × (4–)6–12(–13) mm, juicy throughout; floral remnant persistent. |
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Seeds | reddish brown, sometimes yellowish when fresh, 1–1.2 mm, deeply pitted; testa thin, relatively flexible; anticlinal cell walls sinuate, interstices narrower than pit diameters; pits cavernous or deeply concave. |
chocolate brown with darker reticulation, 1.3 × 0.9 mm, pitted; testa hard; anticlinal cell walls straight; interstices conspicuously narrower than pit diameters; pits bowl-shaped. |
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2n | = 22. |
= 22. |
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Mammillaria heyderi |
Mammillaria viridiflora |
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Phenology | Flowering spring; fruiting fall. | |||||
Habitat | Semidesert grasslands, interior chaparral, pinyon-juniper and oak woodlands, crevices, boulders, canyon sides and gravelly igneous substrates | |||||
Elevation | (800-)1400-2000 m ((2600-)4600-6600 ft) | |||||
Distribution |
AZ; NM; OK; TX; n Mexico
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AZ; NM |
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Discussion | Varieties 2 (2 in the flora). Green fruits of Mammillaria heyderi with fully mature, viable seeds precede the ripe (elongate) fruits by six months to a year. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Mammillaria viridiflora is closely related to M. barbata Engelmann of Mexico, which has priority if the species are combined. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 4, p. 255. | FNA vol. 4, p. 251. | ||||
Parent taxa | Cactaceae > subfam. Cactoideae > Mammillaria | Cactaceae > subfam. Cactoideae > Mammillaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Neomammillaria viridiflora, M. orestera | |||||
Name authority | Muehlenpfordt: Allg. Gartenzeitung 16: 20. (1848) | (Britton & Rose) Boedeker: Mamm.-vergl.-Schlüssel, 36. (1933) | ||||
Web links |