Mammillaria heyderi |
Mammillaria meiacantha |
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biznaga China, Heyder's pincushion cactus, little nipple cactus |
little nipple cactus, nipple cactus |
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Habit | Plants unbranched, protruding relatively little above soil. | Plants unbranched. | ||||
Roots | obconic taproots; secondary roots diffuse. |
short, obconic taproots; secondary roots diffuse. |
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Stems | top-shaped, flat-topped (aerial part sometimes hemispheric in old age or in dense subtropical vegetation), protruding above ground 0–2 × (4–)7.5–15 cm, firm; tubercles 9–15(–20) × 3–7 mm; axils with short wool, bristles absent; cortex and pith not mucilaginous; latex abundant in healthy tissue throughout cortex of stem, tubercles, and sometimes flower receptacle, sticky, white. |
flat-topped (in old age or under dense brush aerial portion of stem hemispheric), 10 × 8–10(–30) cm, firm; tubercles 8–17 × 4–11 mm; axils bearing (at least seasonally) woolly tufts, wool 3–5 mm, bristles absent; cortex and pith not mucilaginous; latex abundant in healthy tissue throughout cortex of stem, tubercles, and sometimes flower receptacle, sticky, white. |
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Spines | (8–)10–18(–27) per areole, usually brownish, darker at tip, glabrous; radial spines (8–)10–22(–26) per areole, white to white-and-brown or brown, needlelike, 6–15(–16) mm, stiff, abaxial spines longest; central spines (0–)1(–4) per areole, porrect or ascending, not hooked, (0.5–)2–8 × 0.15–0.45 mm; subcentral spines 0. |
(6–)7–8(–10) per areole, white, reddish brown, gray, or yellowish, glabrous; radial spines (5–)6–7(–9) per areole, largest spines reddish brown, gray, or yellowish, tips blackish or dark brown, needlelike, 6.5–13.5 × 0.3–0.6 mm, stiff; central spines (0–)1 per areole, usually ascending and inconspicuous against radial spines, sometimes porrect, straight or slightly curved, largest spines (3–)5–12 × 0.3–0.7 mm; subcentral spines 0. |
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Flowers | 1.9–3.8 × 1.5–3 cm; outermost tepal margins entire; inner tepals white, greenish or cream to pale pink, with tan, pink, greenish, or brownish midstripes, 11–19 × 2–2.5 mm; stigma lobes externally green, internally green or red (or pink), 2.5–3 mm. |
2.5–3.5 × 1.9–3.5(–4.4) cm; outermost tepal margins entire; inner tepals white to pale pink, often with pink or lavender midstripes, 8–15 mm; stigma lobes light green, 3–5 mm. |
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Fruits | brilliant red: scarlet, carmine, or crimson, obovoid to clavate, 10–35(–40) × 5–8 mm, juicy only in fruit walls; floral remnant weakly persistent. |
purplish pink, clavate to obovoid, 20–32 mm, juicy only in fruit walls; floral remnant weakly persistent. |
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Seeds | reddish brown, sometimes yellowish when fresh, 1–1.2 mm, deeply pitted; testa thin, relatively flexible; anticlinal cell walls sinuate, interstices narrower than pit diameters; pits cavernous or deeply concave. |
reddish brown, 1.1–1.2 mm; pitted; testa leathery to hard, anticlinal cell walls strongly undulate, interstices much narrower than pit diam., pits deeply concave, elongate. |
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2n | = 22. |
= 22. |
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Mammillaria heyderi |
Mammillaria meiacantha |
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Phenology | Flowering Mar–May(-Jun); fruiting Oct–Mar. | |||||
Habitat | Great Plains grasslands, pine-oak woodlands, ecotone between Chihuahuan desert scrub adjacent more mesic habitats | |||||
Elevation | 900-2500 m (3000-8200 ft) | |||||
Distribution |
AZ; NM; OK; TX; n Mexico
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NM; TX; Mexico |
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Discussion | Varieties 2 (2 in the flora). Green fruits of Mammillaria heyderi with fully mature, viable seeds precede the ripe (elongate) fruits by six months to a year. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Mammillaria meiacantha is regionally sympatric with both varieties of M. heyderi, without resulting in hybridization. West of the Rio Grande, reports of M. meiacantha probably are misidentifications of M. heyderi var. bullingtoniana. Mammillaria meiacantha has not been documented for Arizona. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 4, p. 255. | FNA vol. 4, p. 256. | ||||
Parent taxa | Cactaceae > subfam. Cactoideae > Mammillaria | Cactaceae > subfam. Cactoideae > Mammillaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | M. gummifera var. meiacantha, M. heyderi var. meiacantha, M. runyonii | |||||
Name authority | Muehlenpfordt: Allg. Gartenzeitung 16: 20. (1848) | Engelmann: Proc. Amer. Acad. Arts 3: 263. (1856) | ||||
Web links |