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biznaga China, Heyder's pincushion cactus, little nipple cactus

little nipple cactus, nipple cactus

Habit Plants unbranched, protruding relatively little above soil. Plants unbranched.
Roots

obconic taproots;

secondary roots diffuse.

short, obconic taproots;

secondary roots diffuse.

Stems

top-shaped, flat-topped (aerial part sometimes hemispheric in old age or in dense subtropical vegetation), protruding above ground 0–2 × (4–)7.5–15 cm, firm;

tubercles 9–15(–20) × 3–7 mm;

axils with short wool, bristles absent;

cortex and pith not mucilaginous;

latex abundant in healthy tissue throughout cortex of stem, tubercles, and sometimes flower receptacle, sticky, white.

flat-topped (in old age or under dense brush aerial portion of stem hemispheric), 10 × 8–10(–30) cm, firm;

tubercles 8–17 × 4–11 mm;

axils bearing (at least seasonally) woolly tufts, wool 3–5 mm, bristles absent;

cortex and pith not mucilaginous;

latex abundant in healthy tissue throughout cortex of stem, tubercles, and sometimes flower receptacle, sticky, white.

Spines

(8–)10–18(–27) per areole, usually brownish, darker at tip, glabrous;

radial spines (8–)10–22(–26) per areole, white to white-and-brown or brown, needlelike, 6–15(–16) mm, stiff, abaxial spines longest;

central spines (0–)1(–4) per areole, porrect or ascending, not hooked, (0.5–)2–8 × 0.15–0.45 mm;

subcentral spines 0.

(6–)7–8(–10) per areole, white, reddish brown, gray, or yellowish, glabrous;

radial spines (5–)6–7(–9) per areole, largest spines reddish brown, gray, or yellowish, tips blackish or dark brown, needlelike, 6.5–13.5 × 0.3–0.6 mm, stiff;

central spines (0–)1 per areole, usually ascending and inconspicuous against radial spines, sometimes porrect, straight or slightly curved, largest spines (3–)5–12 × 0.3–0.7 mm;

subcentral spines 0.

Flowers

1.9–3.8 × 1.5–3 cm;

outermost tepal margins entire;

inner tepals white, greenish or cream to pale pink, with tan, pink, greenish, or brownish midstripes, 11–19 × 2–2.5 mm;

stigma lobes externally green, internally green or red (or pink), 2.5–3 mm.

2.5–3.5 × 1.9–3.5(–4.4) cm;

outermost tepal margins entire;

inner tepals white to pale pink, often with pink or lavender midstripes, 8–15 mm;

stigma lobes light green, 3–5 mm.

Fruits

brilliant red: scarlet, carmine, or crimson, obovoid to clavate, 10–35(–40) × 5–8 mm, juicy only in fruit walls;

floral remnant weakly persistent.

purplish pink, clavate to obovoid, 20–32 mm, juicy only in fruit walls;

floral remnant weakly persistent.

Seeds

reddish brown, sometimes yellowish when fresh, 1–1.2 mm, deeply pitted;

testa thin, relatively flexible;

anticlinal cell walls sinuate, interstices narrower than pit diameters;

pits cavernous or deeply concave.

reddish brown, 1.1–1.2 mm; pitted;

testa leathery to hard, anticlinal cell walls strongly undulate, interstices much narrower than pit diam., pits deeply concave, elongate.

2n

= 22.

= 22.

Mammillaria heyderi

Mammillaria meiacantha

Phenology Flowering Mar–May(-Jun); fruiting Oct–Mar.
Habitat Great Plains grasslands, pine-oak woodlands, ecotone between Chihuahuan desert scrub adjacent more mesic habitats
Elevation 900-2500 m (3000-8200 ft)
Distribution
from FNA
AZ; NM; OK; TX; n Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
NM; TX; Mexico
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Green fruits of Mammillaria heyderi with fully mature, viable seeds precede the ripe (elongate) fruits by six months to a year.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Mammillaria meiacantha is regionally sympatric with both varieties of M. heyderi, without resulting in hybridization. West of the Rio Grande, reports of M. meiacantha probably are misidentifications of M. heyderi var. bullingtoniana. Mammillaria meiacantha has not been documented for Arizona.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Central spines 0.15-0.35 mm diam.; radial spines (7-)13-17(-26) per areole, abaxial radial spines 6-11(-16) mm; e of El Paso, Texas, New Mexico, Oklahoma.
var. heyderi
1. Central spines 0.35-0.45 mm diam.; radial spines 10-14 per areole, abaxial radial spines 9-15 mm; El Paso, Texas, w to Arizona
var. bullingtoniana
Source FNA vol. 4, p. 255. FNA vol. 4, p. 256.
Parent taxa Cactaceae > subfam. Cactoideae > Mammillaria Cactaceae > subfam. Cactoideae > Mammillaria
Sibling taxa
M. dioica, M. grahamii, M. lasiacantha, M. macdougalii, M. mainiae, M. meiacantha, M. pottsii, M. prolifera, M. sphaerica, M. tetrancistra, M. thornberi, M. viridiflora, M. wrightii
M. dioica, M. grahamii, M. heyderi, M. lasiacantha, M. macdougalii, M. mainiae, M. pottsii, M. prolifera, M. sphaerica, M. tetrancistra, M. thornberi, M. viridiflora, M. wrightii
Subordinate taxa
M. heyderi var. bullingtoniana, M. heyderi var. heyderi
Synonyms M. gummifera var. meiacantha, M. heyderi var. meiacantha, M. runyonii
Name authority Muehlenpfordt: Allg. Gartenzeitung 16: 20. (1848) Engelmann: Proc. Amer. Acad. Arts 3: 263. (1856)
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