Mammillaria heyderi |
Mammillaria macdougalii |
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biznaga China, Heyder's pincushion cactus, little nipple cactus |
cream cactus |
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Habit | Plants unbranched, protruding relatively little above soil. | Plants unbranched, deeply seated in substrate. | ||||
Roots | obconic taproots; secondary roots diffuse. |
short, obconic taproots; secondary roots diffuse. |
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Stems | top-shaped, flat-topped (aerial part sometimes hemispheric in old age or in dense subtropical vegetation), protruding above ground 0–2 × (4–)7.5–15 cm, firm; tubercles 9–15(–20) × 3–7 mm; axils with short wool, bristles absent; cortex and pith not mucilaginous; latex abundant in healthy tissue throughout cortex of stem, tubercles, and sometimes flower receptacle, sticky, white. |
usually flat-topped (in old age or under dense brush hemispheric), 4–20 × 9-25(–30) cm, firm; tubercles 10–17 × 6–8.5 mm; axils woolly, wool 5–8 mm, bristles absent; cortex and pith not mucilaginous; latex sticky white, abundant in healthy tissue throughout cortex of stem, tubercles, and sometimes flower receptacles. |
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Spines | (8–)10–18(–27) per areole, usually brownish, darker at tip, glabrous; radial spines (8–)10–22(–26) per areole, white to white-and-brown or brown, needlelike, 6–15(–16) mm, stiff, abaxial spines longest; central spines (0–)1(–4) per areole, porrect or ascending, not hooked, (0.5–)2–8 × 0.15–0.45 mm; subcentral spines 0. |
11–14(–16) per areole, white to straw yellow, glabrous; radial spines 10–12 per areole, whitish to pale yellowish tan, needlelike, 20 mm, stiff; central spines 1–2(–4) per areole, ± porrect or 1 ascending and 1 descending, straight or slightly curved, 10 mm; subcentral spines 0. |
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Flowers | 1.9–3.8 × 1.5–3 cm; outermost tepal margins entire; inner tepals white, greenish or cream to pale pink, with tan, pink, greenish, or brownish midstripes, 11–19 × 2–2.5 mm; stigma lobes externally green, internally green or red (or pink), 2.5–3 mm. |
3–4 × 2.5–3.5 cm; outermost tepal margins densely minutely fimbriate (at 10x); inner tepals greenish white to greenish yellow, 14–17(–25) × 2–3 mm; stigma lobes yellow-green to pale green, 2–3 mm. |
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Fruits | brilliant red: scarlet, carmine, or crimson, obovoid to clavate, 10–35(–40) × 5–8 mm, juicy only in fruit walls; floral remnant weakly persistent. |
whitish green proximally, pale green distally with pale rose-purple on sunlit portions, obovoid to clavate, 15–26 × 7.5–15 mm, juicy only in fruit walls; floral remnant persistent. |
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Seeds | reddish brown, sometimes yellowish when fresh, 1–1.2 mm, deeply pitted; testa thin, relatively flexible; anticlinal cell walls sinuate, interstices narrower than pit diameters; pits cavernous or deeply concave. |
reddish brown or orange, 1.1–1.3 × 0.9 × 0.8 mm, pitted; testa intermediate, anticlinal cell walls undulate, interstices much narrower than pit diam., pits deeply concave, elongate. |
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2n | = 22. |
= 22. |
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Mammillaria heyderi |
Mammillaria macdougalii |
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Phenology | Flowering Mar–May; fruiting Oct–Mar. | |||||
Habitat | Rocky slopes and ridges, ecotone between Sonoran desert scrub and chaparral, oak woodlands, grasslands | |||||
Elevation | 1100-1800 m (3600-5900 ft) | |||||
Distribution |
AZ; NM; OK; TX; n Mexico
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AZ; Mexico (Sonora) |
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Discussion | Varieties 2 (2 in the flora). Green fruits of Mammillaria heyderi with fully mature, viable seeds precede the ripe (elongate) fruits by six months to a year. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Mammillaria macdougalii is the characteristic flat-topped mammillaria of the mountains near Tucson and Nogales, Arizona. Records from farther east are all (insofar as can be determined) misidentifications of M. heyderi var. bullingtoniana. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 4, p. 255. | FNA vol. 4, p. 257. | ||||
Parent taxa | Cactaceae > subfam. Cactoideae > Mammillaria | Cactaceae > subfam. Cactoideae > Mammillaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | M. gummifera var. macdougalii, M. heyderi var. macdougalii | |||||
Name authority | Muehlenpfordt: Allg. Gartenzeitung 16: 20. (1848) | Rose: in L. H. Bailey, Stand. Cycl. Hort. 4: 1982. (1916) | ||||
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