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biznaga China, Heyder's pincushion cactus, little nipple cactus

cream cactus

Habit Plants unbranched, protruding relatively little above soil. Plants unbranched, deeply seated in substrate.
Roots

obconic taproots;

secondary roots diffuse.

short, obconic taproots;

secondary roots diffuse.

Stems

top-shaped, flat-topped (aerial part sometimes hemispheric in old age or in dense subtropical vegetation), protruding above ground 0–2 × (4–)7.5–15 cm, firm;

tubercles 9–15(–20) × 3–7 mm;

axils with short wool, bristles absent;

cortex and pith not mucilaginous;

latex abundant in healthy tissue throughout cortex of stem, tubercles, and sometimes flower receptacle, sticky, white.

usually flat-topped (in old age or under dense brush hemispheric), 4–20 × 9-25(–30) cm, firm;

tubercles 10–17 × 6–8.5 mm;

axils woolly, wool 5–8 mm, bristles absent;

cortex and pith not mucilaginous;

latex sticky white, abundant in healthy tissue throughout cortex of stem, tubercles, and sometimes flower receptacles.

Spines

(8–)10–18(–27) per areole, usually brownish, darker at tip, glabrous;

radial spines (8–)10–22(–26) per areole, white to white-and-brown or brown, needlelike, 6–15(–16) mm, stiff, abaxial spines longest;

central spines (0–)1(–4) per areole, porrect or ascending, not hooked, (0.5–)2–8 × 0.15–0.45 mm;

subcentral spines 0.

11–14(–16) per areole, white to straw yellow, glabrous;

radial spines 10–12 per areole, whitish to pale yellowish tan, needlelike, 20 mm, stiff;

central spines 1–2(–4) per areole, ± porrect or 1 ascending and 1 descending, straight or slightly curved, 10 mm;

subcentral spines 0.

Flowers

1.9–3.8 × 1.5–3 cm;

outermost tepal margins entire;

inner tepals white, greenish or cream to pale pink, with tan, pink, greenish, or brownish midstripes, 11–19 × 2–2.5 mm;

stigma lobes externally green, internally green or red (or pink), 2.5–3 mm.

3–4 × 2.5–3.5 cm;

outermost tepal margins densely minutely fimbriate (at 10x);

inner tepals greenish white to greenish yellow, 14–17(–25) × 2–3 mm;

stigma lobes yellow-green to pale green, 2–3 mm.

Fruits

brilliant red: scarlet, carmine, or crimson, obovoid to clavate, 10–35(–40) × 5–8 mm, juicy only in fruit walls;

floral remnant weakly persistent.

whitish green proximally, pale green distally with pale rose-purple on sunlit portions, obovoid to clavate, 15–26 × 7.5–15 mm, juicy only in fruit walls;

floral remnant persistent.

Seeds

reddish brown, sometimes yellowish when fresh, 1–1.2 mm, deeply pitted;

testa thin, relatively flexible;

anticlinal cell walls sinuate, interstices narrower than pit diameters;

pits cavernous or deeply concave.

reddish brown or orange, 1.1–1.3 × 0.9 × 0.8 mm, pitted;

testa intermediate, anticlinal cell walls undulate, interstices much narrower than pit diam., pits deeply concave, elongate.

2n

= 22.

= 22.

Mammillaria heyderi

Mammillaria macdougalii

Phenology Flowering Mar–May; fruiting Oct–Mar.
Habitat Rocky slopes and ridges, ecotone between Sonoran desert scrub and chaparral, oak woodlands, grasslands
Elevation 1100-1800 m (3600-5900 ft)
Distribution
from FNA
AZ; NM; OK; TX; n Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; Mexico (Sonora)
Discussion

Varieties 2 (2 in the flora).

Green fruits of Mammillaria heyderi with fully mature, viable seeds precede the ripe (elongate) fruits by six months to a year.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Mammillaria macdougalii is the characteristic flat-topped mammillaria of the mountains near Tucson and Nogales, Arizona. Records from farther east are all (insofar as can be determined) misidentifications of M. heyderi var. bullingtoniana.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Central spines 0.15-0.35 mm diam.; radial spines (7-)13-17(-26) per areole, abaxial radial spines 6-11(-16) mm; e of El Paso, Texas, New Mexico, Oklahoma.
var. heyderi
1. Central spines 0.35-0.45 mm diam.; radial spines 10-14 per areole, abaxial radial spines 9-15 mm; El Paso, Texas, w to Arizona
var. bullingtoniana
Source FNA vol. 4, p. 255. FNA vol. 4, p. 257.
Parent taxa Cactaceae > subfam. Cactoideae > Mammillaria Cactaceae > subfam. Cactoideae > Mammillaria
Sibling taxa
M. dioica, M. grahamii, M. lasiacantha, M. macdougalii, M. mainiae, M. meiacantha, M. pottsii, M. prolifera, M. sphaerica, M. tetrancistra, M. thornberi, M. viridiflora, M. wrightii
M. dioica, M. grahamii, M. heyderi, M. lasiacantha, M. mainiae, M. meiacantha, M. pottsii, M. prolifera, M. sphaerica, M. tetrancistra, M. thornberi, M. viridiflora, M. wrightii
Subordinate taxa
M. heyderi var. bullingtoniana, M. heyderi var. heyderi
Synonyms M. gummifera var. macdougalii, M. heyderi var. macdougalii
Name authority Muehlenpfordt: Allg. Gartenzeitung 16: 20. (1848) Rose: in L. H. Bailey, Stand. Cycl. Hort. 4: 1982. (1916)
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