Mammillaria heyderi |
Mammillaria grahamii |
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biznaga China, Heyder's pincushion cactus, little nipple cactus |
Arizona fishhook cactus, Graham' s nipple cactus, Graham's fishhook cactus, Graham's pincushion cactus |
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Habit | Plants unbranched, protruding relatively little above soil. | Plants branched or unbranched; branches 0–9(–17). | ||||
Roots | obconic taproots; secondary roots diffuse. |
diffuse; upper portion not enlarged. |
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Stems | top-shaped, flat-topped (aerial part sometimes hemispheric in old age or in dense subtropical vegetation), protruding above ground 0–2 × (4–)7.5–15 cm, firm; tubercles 9–15(–20) × 3–7 mm; axils with short wool, bristles absent; cortex and pith not mucilaginous; latex abundant in healthy tissue throughout cortex of stem, tubercles, and sometimes flower receptacle, sticky, white. |
spheric to cylindric, usually (4–)5–16(–30) × (2.3–)3.5–6.8 cm, firm; tubercles 4.8–12(–15) × 3.5–7 mm; axils appearing naked; cortex and pith not mucilaginous; latex absent. |
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Spines | (8–)10–18(–27) per areole, usually brownish, darker at tip, glabrous; radial spines (8–)10–22(–26) per areole, white to white-and-brown or brown, needlelike, 6–15(–16) mm, stiff, abaxial spines longest; central spines (0–)1(–4) per areole, porrect or ascending, not hooked, (0.5–)2–8 × 0.15–0.45 mm; subcentral spines 0. |
(19–)26–33(–38) per areole, glabrous; radial spines 17–35 per areole whitish or pale tan, bristlelike, 6–12 × 0.1–0.15 mm, stiff; central spines (2–)3–4 per areole, 1–3(–4) spines at least hooked (uncommonly 0 hooked), reddish to purplish brown to almost black (rarely golden brown), abaxial 1 porrect, others inconspicuous, appressed against radial spines, hookless porrect spines (3–)9.5–25 × 0.1–0.5 mm; subcentral spines 1–3 per areole, adaxial to central spines, sometimes transitional to central spines, usually straight and barely distinguishable from radial spines. |
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Flowers | 1.9–3.8 × 1.5–3 cm; outermost tepal margins entire; inner tepals white, greenish or cream to pale pink, with tan, pink, greenish, or brownish midstripes, 11–19 × 2–2.5 mm; stigma lobes externally green, internally green or red (or pink), 2.5–3 mm. |
ca. 2 × 1.8–3.5(–4.5) cm; outermost tepal margins minutely fringed; inner tepals bright rose-pink or rose-purple, 10–16 × 4–8 mm; stigma lobes yellow-green to green, 3–7 mm. |
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Fruits | brilliant red: scarlet, carmine, or crimson, obovoid to clavate, 10–35(–40) × 5–8 mm, juicy only in fruit walls; floral remnant weakly persistent. |
green, turning bright red, scarlet, or carmine (rarely yellowish), elongating until clavate after color change to red is complete, 12–29 × 5–8 mm, juicy only in fruit walls; floral remnant persistent. |
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Seeds | reddish brown, sometimes yellowish when fresh, 1–1.2 mm, deeply pitted; testa thin, relatively flexible; anticlinal cell walls sinuate, interstices narrower than pit diameters; pits cavernous or deeply concave. |
black, 0.8–1 × 0.7–0.9 mm, pitted; testa hard, brittle; anticlinal cell walls straight; interstices conspicuously wider than pit diameters; pits bowl-shaped. |
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2n | = 22. |
= 22. |
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Mammillaria heyderi |
Mammillaria grahamii |
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Phenology | Flowering Apr–Sep; fruiting Sep–Mar. | |||||
Habitat | Chihuahuan and Sonoran desert scrub, grasslands, interior chaparral, oak woodlands, alluvial slopes, hills, canyons, silty, sandy, gravelly, or rocky soils of igneous or calcareous origin | |||||
Elevation | 80-1400 m (300-4600 ft) | |||||
Distribution |
AZ; NM; OK; TX; n Mexico
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AZ; CA; NM; TX; Mexico (Chihuahua, Sonora)
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Discussion | Varieties 2 (2 in the flora). Green fruits of Mammillaria heyderi with fully mature, viable seeds precede the ripe (elongate) fruits by six months to a year. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Mammillaria grahamii is geographically variable. Past attempts to distinguish larger or western individuals as M. microcarpa have proven arbitrary. All spine hooks on a plant may be oriented in the same direction, a trait sometimes mistakenly said to be limited to Mammillaria mainiae. Plants with short, straight central spines (rarely a mixture of both hooked and straight spines on the same stem) occur in some populations; they have been called M. oliviae or M. grahamii var. oliviae. The name Mammillaria microcarpa Engelmann has been widely used but was not validly published until after M. grahamii. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 4, p. 255. | FNA vol. 4. | ||||
Parent taxa | Cactaceae > subfam. Cactoideae > Mammillaria | Cactaceae > subfam. Cactoideae > Mammillaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | M. grahamii var. oliviae, M. microcarpa, Neomammillaria milleri | |||||
Name authority | Muehlenpfordt: Allg. Gartenzeitung 16: 20. (1848) | Engelmann: Proc. Amer. Acad. Arts 3: 262. (1856) | ||||
Web links |