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biznaga China, Heyder's pincushion cactus, little nipple cactus

Habit Plants unbranched, protruding relatively little above soil.
Roots

obconic taproots;

secondary roots diffuse.

Stems

top-shaped, flat-topped (aerial part sometimes hemispheric in old age or in dense subtropical vegetation), protruding above ground 0–2 × (4–)7.5–15 cm, firm;

tubercles 9–15(–20) × 3–7 mm;

axils with short wool, bristles absent;

cortex and pith not mucilaginous;

latex abundant in healthy tissue throughout cortex of stem, tubercles, and sometimes flower receptacle, sticky, white.

Spines

(8–)10–18(–27) per areole, usually brownish, darker at tip, glabrous;

radial spines (8–)10–22(–26) per areole, white to white-and-brown or brown, needlelike, 6–15(–16) mm, stiff, abaxial spines longest;

central spines (0–)1(–4) per areole, porrect or ascending, not hooked, (0.5–)2–8 × 0.15–0.45 mm;

subcentral spines 0.

Flowers

1.9–3.8 × 1.5–3 cm;

outermost tepal margins entire;

inner tepals white, greenish or cream to pale pink, with tan, pink, greenish, or brownish midstripes, 11–19 × 2–2.5 mm;

stigma lobes externally green, internally green or red (or pink), 2.5–3 mm.

Fruits

brilliant red: scarlet, carmine, or crimson, obovoid to clavate, 10–35(–40) × 5–8 mm, juicy only in fruit walls;

floral remnant weakly persistent.

Seeds

reddish brown, sometimes yellowish when fresh, 1–1.2 mm, deeply pitted;

testa thin, relatively flexible;

anticlinal cell walls sinuate, interstices narrower than pit diameters;

pits cavernous or deeply concave.

2n

= 22.

Mammillaria heyderi

Distribution
from FNA
AZ; NM; OK; TX; n Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Green fruits of Mammillaria heyderi with fully mature, viable seeds precede the ripe (elongate) fruits by six months to a year.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Central spines 0.15-0.35 mm diam.; radial spines (7-)13-17(-26) per areole, abaxial radial spines 6-11(-16) mm; e of El Paso, Texas, New Mexico, Oklahoma.
var. heyderi
1. Central spines 0.35-0.45 mm diam.; radial spines 10-14 per areole, abaxial radial spines 9-15 mm; El Paso, Texas, w to Arizona
var. bullingtoniana
Source FNA vol. 4, p. 255.
Parent taxa Cactaceae > subfam. Cactoideae > Mammillaria
Sibling taxa
M. dioica, M. grahamii, M. lasiacantha, M. macdougalii, M. mainiae, M. meiacantha, M. pottsii, M. prolifera, M. sphaerica, M. tetrancistra, M. thornberi, M. viridiflora, M. wrightii
Subordinate taxa
M. heyderi var. bullingtoniana, M. heyderi var. heyderi
Name authority Muehlenpfordt: Allg. Gartenzeitung 16: 20. (1848)
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