Mammillaria dioica |
Mammillaria heyderi |
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California fishhook cactus, fish hook cactus, strawberry cactus |
biznaga China, Heyder's pincushion cactus, little nipple cactus |
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Habit | Plants unbranched or branched; branches 0–50. | Plants unbranched, protruding relatively little above soil. | ||||
Roots | diffuse, upper portion not enlarged. |
obconic taproots; secondary roots diffuse. |
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Stems | nearly spheric to more often cylindric or long cylindric, 5–30 × 5–7 cm, firm; tubercles 5–12 × 3–7 mm; axils woolly, bearing 4–15 bristles (0 in young growth) as long as tubercles; cortex and pith not mucilaginous; latex absent. |
top-shaped, flat-topped (aerial part sometimes hemispheric in old age or in dense subtropical vegetation), protruding above ground 0–2 × (4–)7.5–15 cm, firm; tubercles 9–15(–20) × 3–7 mm; axils with short wool, bristles absent; cortex and pith not mucilaginous; latex abundant in healthy tissue throughout cortex of stem, tubercles, and sometimes flower receptacle, sticky, white. |
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Spines | 14–26 per areole, pinkish or reddish brown to black, glabrous; radial spines 11–22 per areole, usually white, bristlelike, 5–7 mm, stiff; central spines (1–)3–4 per areole, abaxial 1 porrect, hooked, longer, stouter, adaxial central spines ascending with radial spines; subcentral spines 0. |
(8–)10–18(–27) per areole, usually brownish, darker at tip, glabrous; radial spines (8–)10–22(–26) per areole, white to white-and-brown or brown, needlelike, 6–15(–16) mm, stiff, abaxial spines longest; central spines (0–)1(–4) per areole, porrect or ascending, not hooked, (0.5–)2–8 × 0.15–0.45 mm; subcentral spines 0. |
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Flowers | 10–22 mm; outermost tepals entire or short fringed; inner tepals cream, usually with pinkish or reddish midstripes, longer in bisexual flowers, 5.4 mm diam.; stigma lobes yellow to greenish yellow or brownish green, 8 mm. |
1.9–3.8 × 1.5–3 cm; outermost tepal margins entire; inner tepals white, greenish or cream to pale pink, with tan, pink, greenish, or brownish midstripes, 11–19 × 2–2.5 mm; stigma lobes externally green, internally green or red (or pink), 2.5–3 mm. |
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Fruits | bright scarlet, clavate or ovoid, 10–25(–35) × 10 mm, juicy only in fruit walls; floral remnant persistent. |
brilliant red: scarlet, carmine, or crimson, obovoid to clavate, 10–35(–40) × 5–8 mm, juicy only in fruit walls; floral remnant weakly persistent. |
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Seeds | black, 0.8 × 0.6 mm, pitted; testa hard; anticlinal cell walls straight (not undulate); interstices conspicuously wider than pit diameters; pits bowl-shaped. |
reddish brown, sometimes yellowish when fresh, 1–1.2 mm, deeply pitted; testa thin, relatively flexible; anticlinal cell walls sinuate, interstices narrower than pit diameters; pits cavernous or deeply concave. |
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2n | = 66. |
= 22. |
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Mammillaria dioica |
Mammillaria heyderi |
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Phenology | Flowering spring (Mar–May); fruiting summer. | |||||
Habitat | California coastal scrub, Colorado subdivision of Sonoran desert scrub, rocky slopes | |||||
Elevation | 10-1500 m (0-4900 ft) | |||||
Distribution |
CA; Mexico (Baja California)
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AZ; NM; OK; TX; n Mexico
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Discussion | In an inland population in California, Mammillaria dioica was found to be functionally gynodioecious (F. R. Ganders and H. Kennedy 1978), with flowers of some plants bisexual while those of other individuals bear only functionally female flowers with sterile anthers. Coastal populations of the species were not studied and might be “trioecious” with staminate, pistillate, and bisexual flowers on different plants (B. D. Parfitt 1985). Plants of Mammillaria dioica in Mexico are both tetraploid and hexaploid (M. A. T. Johnson 1978). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). Green fruits of Mammillaria heyderi with fully mature, viable seeds precede the ripe (elongate) fruits by six months to a year. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 4, p. 253. | FNA vol. 4, p. 255. | ||||
Parent taxa | Cactaceae > subfam. Cactoideae > Mammillaria | Cactaceae > subfam. Cactoideae > Mammillaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Name authority | K. Brandegee: Erythea 5: 115. (1897) | Muehlenpfordt: Allg. Gartenzeitung 16: 20. (1848) | ||||
Web links |