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Fendler's desert dandelion

desertdandelion, malacothrix

Habit Annuals, 3–15(–25+) cm. Annuals or perennials, 2–70(–200) cm; taprooted (rhizomatous or taproots becoming caudices in M. saxatilis).
Stems

(1–)3–8, ± decumbent or spreading-ascending, branched proximally and distally, glaucous or glabrous.

1–15 (usually from basal rosettes), usually erect, sometimes ± prostrate, usually branched (scapiform in M. californica), usually glabrous (sometimes piloso-hirsute, stipitate-glandular, or tomentose to arachnose or puberulent, at least proximally or in leaf axils).

Leaves

usually basal and cauline;

sessile;

blades mostly oblong or lanceolate to obovate, oblanceolate, or spatulate (often pinnately lobed), ultimate margins entire or ± dentate (faces usually glabrous, sometimes piloso-hirsute or tomentose to arachnose or puberulent).

Cauline leaves

proximal elliptic to oblong-oblanceolate, sometimes pinnately lobed (lobes 2–4+ pairs, oblong to triangular, unequal, apices acute), not fleshy, ultimate margins usually dentate, faces glabrous;

distal reduced (narrowly triangular to linear or filiform, margins dentate or entire).

Peduncles

not inflated distally, usually bracteate.

Involucres

± campanulate, 7–10 × 5–6+ mm.

usually broadly to narrowly campanulate, sometimes hemispheric, (5–22+ ×) 2–22+ mm diam.

Receptacles

not bristly.

flat to ± convex, pitted or smooth, sometimes bristly, epaleate.

Florets

16–88;

corollas yellow (usually with red or purplish abaxial stripes), 6–14 mm;

outer ligules exserted 5–8 mm.

15–270;

corollas yellow or white (sometimes reddish or lavender abaxially; outer ligules exserted 1–15 mm).

Phyllaries

13–25+ in 2–3 series, lance-oblong or lanceolate to lance-linear, subequal, hyaline margins 0.05–0.3 mm wide, faces glabrous.

either (without calyculi) 25–80+ in 4–6+ series and orbiculate to oblong, lance-oblong, lanceolate, or linear, unequal, or (with calyculi) 12–25+ in 2–3 series and oblong or lanceolate to linear, subequal;

margins ± hyaline, 0.05–2.5 mm, apices obtuse to acute or acuminate.

Calyculi

of 5–12, ovate to lanceolate bractlets, hyaline margins 0.05–0.2 mm wide.

0 (outer phyllaries intergrading with inner) or of 3–30+, ± deltate or lanceolate to linear or subulate, subequal to unequal bractlets (in 1–2 series distinct from phyllaries, margins usually hyaline, faces usually glabrous, sometimes arachnose, rarely stipitate-glandular).

Heads

usually in corymbiform to paniculiform arrays (borne singly in M. californica).

Cypselae

± cylindric, 1.8–2.4 mm (distal 0.3 mm slightly expanded, cupped, smooth), ribs not extending to apices, ± equal;

persistent pappi of 12–15, ± deltate teeth (often hidden within cups at apices of cypselae) plus 1–2 bristles.

(monomorphic) stramineous to brown or purplish brown, ± prismatic or cylindro-fusiform, not beaked, ribs 15 (often 5 more prominent than others, apices of ribs sometimes projecting and forming coronas subtending pappi), usually glabrous (sometimes minutely hirtellous or muriculate);

pappi 0, or (single or double) persistent, whitish, crenate crowns or rings of (1–)8–25+ teeth (mostly 0.05–0.1 mm) plus 0–6, coarse, smooth bristles (setiform scales), all in ± 1 series, subtending (i.e., exterior to) the readily falling, inner (or single and only) pappi of 15–35, basally coherent, white, fine, smooth to barbellulate or (proximally) ± plumose bristles in 1 series (falling all together or in groups).

Pollen

70–100% 3-porate.

x

= 7, 9.

2n

= 14.

Malacothrix fendleri

Malacothrix

Phenology Flowering Mar–Jun.
Habitat Grasslands, pinyon-juniper woodlands, creosote bush associations
Elevation 80–2200 m (300–7200 ft)
Distribution
from FNA
AZ; NM; TX; Mexico (Baja California, Sonora)
[WildflowerSearch map]
[BONAP county map]
from USDA
w United States; nw Mexico [Introduced in South America]
[BONAP county map]
Discussion

Malacothrix fendleri grows in the Sonoran Desert. “San Bernardino Co.” as locality for a specimen from the herbarium of J. G. Lemmon in UC (336493) is evidently an error.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 20 (18 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Perennials (sometimes flowering first year)
→ 2
1. Annuals
→ 3
2. Corollas medium yellow; persistent pappi 0
M. incana
2. Corollas white (usually each with abaxial purple stripe); persistent pappi of fimbriate crowns or 20–25, blunt teeth (0.01–0.05 mm) plus 0 bristles
M. saxatilis
3. Involucres hemispheric; phyllaries orbiculate (outer) to oblong or linear, hyaline margins 1–2.5 mm wide
M. coulteri
3. Involucres usually ± campanulate (to hemispheric in M. sonchoides); bractlets of calyculi and/or phyllaries ovate to lanceolate, linear, or subulate, hyaline margins 0.05–0.3(–1) mm wide
→ 4
4. Stems seldom branched (heads usually borne singly on scapiform peduncles); cypsela ribs: 5 more prominent than others
M. californica
4. Stems branched (heads usually in corymbiform to paniculiform arrays); cypsela ribs ± equal, or 5 more prominent than others
→ 5
5. Cauline leaves not or seldom notably reduced distally (Channel Islands, California)
→ 6
5. Cauline leaves usually reduced distally (M. clevelandii and M. similis are known from Channel Islands, California)
→ 9
6. Persistent pappi usually 0, rarely of 1–2 bristles; pollen 70–100% 3-porate
→ 7
6. Persistent pappi of 15–20 teeth (0.01–0.1 mm) plus 1–2 bristles; pollen 70–100% 4-porate
→ 8
7. Stems erect or ± prostrate (forming mats); proximal cauline leaves usually fleshy (lobes nearly equal, apices obtuse); San Miguel, Santa Cruz, and Santa Rosa islands
M. indecora
7. Stems erect (not forming mats); proximal cauline leaves usually not fleshy (lobes usually unequal, apices acute or obtuse); Anacapa, San Clemente, San Nicolas, and Santa Barbara islands
M. foliosa
8. Calyculi 0; phyllaries 31–49 in 5–6+ series, hyaline margins 0.6–1 mm wide; Anacapa and Santa Cruz islands, California
M. squalida
8. Calyculi of 7–11 bractlets; phyllaries 9–13 in 2–3 series, hyaline margins 0.1–0.2 mm wide; Anacapa Island, California
M. junakii
9. Corollas 4–10 mm; outer ligules exserted 1–4 mm
→ 10
9. Corollas (7–)10–23+ mm; outer ligules exserted 5–15+ mm
→ 14
10. Proximal cauline leaves (pinnately lobed, lobes 3–5 pairs, ± equal, bases white-hairy) ± fleshy; persistent pappi of crenate crowns plus 0 bristles; pollen 70–100% 3-porate.
M. phaeocarpa
10. Proximal cauline leaves (pinnately lobed or not) not fleshy; persistent pappi of 8–24+ teeth plus 1–2 bristles; pollen 3- or 4-porate
→ 11
11. Proximal cauline leaves oblanceolate to lance-linear; corollas pale yellow; cypselae ± cylindro-fusiform or prismatic, 1.2–1.8 mm (ribs extending to apices, 5 more prominent than others); persistent pappi of 14–24+ needlelike teeth plus 1 bristle (mainly California)
→ 12
11. Proximal cauline leaves obovate to narrowly oblanceolate; corollas white or pale yellow; cypselae ± cylindro-fusiform, 1.7–2.3 mm (ribs not extending to apices, ± equal); persistent pappi of 8–18 teeth plus 1–2 bristles (Arizona, California, Nevada)
→ 13
12. Stems branched mostly distally, glabrous; cauline leaves: margins usually dentate; cypselae (stramineous to brown) 1.2–1.8 mm; pollen 3-porate (mean diam. 25 µm)
M. clevelandii
12. Stems branched mostly proximally; cauline leaves: margins usually entire; cypselae 1.4–1.7 mm (purplish brown); pollen 4-porate (mean pollen diam. 30 µm)
M. similis
13. Corollas white or pale yellow; cypselae 1.7–2 mm (bases slightly expanded, distal 0.3 mm smooth); pollen 70–100% 3-porate
M. sonorae
13. Corollas usually yellow, sometimes white; cypselae 1.7–2.3 mm (bases not expanded, distal 0.1–0.2 mm smooth); pollen 70–100% 4-porate
M. stebbinsii
14. Proximal cauline leaves pinnately lobed (lobes 2–6+ pairs, filiform or triangular to oblong, subequal to unequal, apices acute), not fleshy, ultimate margins dentate or entire; cypsela ribs ± equal
→ 15
14. Proximal cauline leaves pinnately lobed (lobes 3–8 pairs, ± oblong to triangular, ± equal, apices obtuse or acute), ± fleshy, ultimate margins usually dentate; cypsela ribs ± equal or 5 more prominent than others
→ 16
15. Proximal cauline leaves usually pinnately lobed (lobes filiform); receptacles bristly; cypselae ± cylindro-fusiform (sometimes weakly 5-angled, ribs extending to apices).
M. glabrata
15. Proximal cauline leaves sometimes pinnately lobed (lobes relatively broad, triangular to deltate); receptacles not bristly; cypselae ± cylindro-fusiform (ribs not extending to apices, distal 0.3 mm of cypselae smooth)
M. fendleri
16. Corollas white or yellow (usually with lavender abaxially); cypselae 1.2–2 mm; persistent pappi 0
M. floccifera
16. Corollas lemon or medium yellow; cypselae 1.8–4 mm; persistent pappi of 12–25+ irregular, blunt teeth plus 0–6 bristles
→ 17
17. Distal cauline leaves narrowly triangular to linear (bases usually ± dilated, ± clasping); persistent pappi of crenate crowns or 12–25+ teeth plus 0 bristles.
M. sonchoides
17. Distal cauline leaves ± elliptic to linear (bases narrowly cuneate); persistent pappi of 12–15 teeth plus 0–6 bristles
M. torreyi
Source FNA vol. 19, p. 314. FNA vol. 19, p. 310. Author: W. S. Davis.
Parent taxa Asteraceae > tribe Cichorieae > Malacothrix Asteraceae > tribe Cichorieae
Sibling taxa
M. californica, M. clevelandii, M. coulteri, M. floccifera, M. foliosa, M. glabrata, M. incana, M. indecora, M. junakii, M. phaeocarpa, M. saxatilis, M. similis, M. sonchoides, M. sonorae, M. squalida, M. stebbinsii, M. torreyi
Subordinate taxa
M. californica, M. clevelandii, M. coulteri, M. fendleri, M. floccifera, M. foliosa, M. glabrata, M. incana, M. indecora, M. junakii, M. phaeocarpa, M. saxatilis, M. similis, M. sonchoides, M. sonorae, M. squalida, M. stebbinsii, M. torreyi
Name authority A. Gray: Smithsonian Contr. Knowl. 5(6): 104. (1853) de Candolle: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 7: 192. (1838)
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