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false serviceberry

Habit Shrubs, 10–30[–50] dm. Shrubs or trees, sometimes subshrubs or herbs.
Stems

1–20, ascending, much branched, often ± twisted, young stems terete, closely villous, smooth;

bark brown or gray, young stems not hairy; long shoots present, short shoots sometimes present; unarmed;

grayish- or whitish-tomentose, sometimes glabrate; overwintering buds tomentose.

Leaves

drought-deciduous or persistent, cauline, simple, slightly conduplicate;

stipules tardily deciduous or ± persistent, free, lanceolate, glandlike, margins entire;

petiole present;

blade elliptic or oblong-ovate to orbiculate, 0.5–1.5[–3.1] cm, leathery, margins flat, denticulate or entire, teeth gland-tipped, venation pinnate, abaxial surface tomentose, adaxial ± glabrous.

alternate, sometimes opposite, simple, sometimes pinnately compound;

stipules present or absent.

Inflorescences

terminal on leafy shoots of season, 1–5[–7]-flowered, corymbs [panicles], axes glabrous or hairy;

bracts absent;

bracteoles sometimes present.

Pedicels

present.

Flowers

perianth and androecium epigynous, 10 mm diam.;

hypanthium campanulate [urceolate], (2–)3–4 mm, distally copiously white-tomentose;

sepals 5, spreading, nearly orbiculate (inner broadly deltate);

petals 5, white, round or kidney-shaped, base short-clawed;

stamens [8–]14–20[–22], longer or shorter than petals;

carpels 2 or 3[–5], barely connate or distinct, adnate to proximal 1/2–4/5 of hypanthium, ovary appearing 4–10-loculed by false partitions, densely hairy, styles 2 or 3, ventral, distinct;

ovules 2 (separated by false partition).

torus absent or minute;

carpels 1–5(–8), distinct or +/- connate (Maleae), free or +/- adnate to hypanthium (many Maleae), styles distinct or +/- connate (some Maleae);

ovules (1 or)2(–5+), collateral, clustered, or biseriate.

Fruits

berrylike pomes, translucent, vivid pink, drying purplish black, globose, 6–10 mm, glabrous;

flesh whitish;

hypanthium persistent;

sepals persistent, erect or recurving;

carpels cartilaginous;

styles persistent.

follicles aggregated or not, capsules, drupes aggregated or not, aggregated drupelets, pomes, or aggregated nutlets, rarely achenes or aggregated achenes;

styles persistent or deciduous, not elongate (elongate in Gillenieae).

Seeds

4–6(–10).

x

= 8, 9, 15, 17.

Malacomeles

Rosaceae subfam. amygdaloideae

Distribution
from USDA
United States; Mexico; Central America
[BONAP county map]
from FNA
HI; North America; Mexico; Central America; South America; Europe; Asia; Africa; Atlantic Islands (Madeira); Australia
Discussion

Species 2 (1 in the flora).

Malacomeles shares with Amelanchier and Peraphyllum the feature of false partitions within the carpels; the partitions partially divide the locules and make the fruit appear to have twice as many locules as the number of carpels (G. N. Jones 1945; K. R. Robertson et al. 1991; J. R. Rohrer et al. 1991). Sequences from multiple chloroplast and nuclear genes indicate that Malacomeles is closely related to Amelanchier and Peraphyllum (C. S. Campbell et al. 2007).

The other species in the genus, Malacomeles nervosa (Decaisne) G. N. Jones, is known from Nuevo León, Mexico south to Guatemala.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cyanogenic glycosides are usually present in Amygdaloideae; sorbitol is present.

The name Amygdaloideae Arnott (1832) has priority over Spiraeoideae Arnott (1832), used by D. Potter et al. (2007), because Amygdalaceae (1820) is an earlier conserved name.

Tribes 9, genera 55, species ca. 1300 (9 tribes, 38 genera, 361 species, including 20 hybrids, in the flora)

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 645. Author: Christopher S. Campbell. FNA vol. 9, p. 345. Author: Luc Brouillet.
Parent taxa Rosaceae > subfam. Amygdaloideae > tribe Maleae Rosaceae
Subordinate taxa
M. denticulata
Synonyms Cotoneaster section M.
Name authority (Decaisne) Decaisne: Ann. Gén. Hort. 23: 156. (1882) Arnott: Botany, 107. (1832)
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