Madia sativa |
Madia gracilis |
|
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Chile tarplant, Chile tarweed, Chilean tarplant, Chilean tarweed, coast tarweed, coastal tarweed |
common tarweed, grassy tarplant, grassy tarweed, gumweed, slender tarweed |
|
Habit | Plants (0.3–)35–100(–240) cm, self-compatible (heads not showy). | Plants 6–100 cm, self-compatible (heads not showy). |
Stems | hirsute and glandular-pubescent, glands yellowish, purple, or black, lateral branches rarely surpassing main stems. |
proximally pilose to hirsute, distally glandular-pubescent, glands yelloish, purple, or black, lateral branches seldom surpassing main stems. |
Leaf | blades broadly lanceolate to linear-oblong or linear, 2–18 cm × 3–18(–29) mm. |
blades oblong to linear, 1–10(–15) cm × 1–8(–10) mm. |
Involucres | ovoid to urceolate, 6–16 mm. |
depressed-globose to urceolate, 5–10 mm. |
Ray florets | (5–)8–13; corollas greenish yellow or sometimes purplish red abaxially or throughout, laminae 1.5–4 mm. |
3–10; corollas lemon yellow or greenish yellow, laminae 1.5–8 mm. |
Disc florets | 11–14, bisexual, fertile; corollas 2–5 mm, pubescent; anthers ± dark purple. |
2–16+, bisexual, fertile; corollas 2.5–5 mm, pubescent; anthers ± dark purple. |
Phyllaries | hirsute and glandular-pubescent, glands yellowish, purple, or black, apices erect or ± reflexed, flat. |
sometimes hirsute, always finely or coarsely glandular-pubescent, glands yellowish, purple, or black, apices erect or ± reflexed, flat. |
Heads | in usually crowded, paniculiform, racemiform, or spiciform arrays. |
in ± open, paniculiform or racemiform arrays. |
Disc cypselae | similar. |
similar. |
Ray cypselae | black or brown, sometimes mottled, dull, compressed, beakless. |
black, purple, or mottled, dull, compressed, beakless (or nearly so). |
Paleae | mostly persistent, connate 1/2+ their lengths. |
mostly persistent, connate 1/2+ their lengths. |
2n | = 32. |
= 32, 48. |
Madia sativa |
Madia gracilis |
|
Phenology | Flowering May–Oct. | Flowering Apr–Aug. |
Habitat | Grasslands, openings in shrublands and woods, disturbed sites, stream banks, roadsides | Open or partially shaded slopes or flats in grasslands, meadows, shrublands, woodlands, and forests, disturbed sites, stream banks, roadsides, coarse to fine textured soils, sometimes serpentine |
Elevation | 0–1000 m (0–3300 ft) | 0–2500 m (0–8200 ft) |
Distribution |
CA; OR; WA; BC; South America (Argentina, Chile) [Pacific Islands (Hawaii, probably introduced)]
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CA; ID; MT; NV; OR; UT; WA; BC; Mexico (Baja California)
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Discussion | In North America, Madia sativa occurs on the Pacific Coast from California to British Columbia, sporadically in coastal ranges, and rarely eastward. Reports of M. sativa from Ontario and Quebec and from Alaska, Connecticut, Georgia, Idaho, Indiana, Iowa, Maine, Maryland, Massachusetts, New York, North Carolina, Pennsylvania, Vermont, and Wisconsin are putative waifs or misidentified M. glomerata. Molecular data and greenhouse studies have indicated that plants referable to M. capitata and M. sativa in California are not distinct (B. G. Baldwin, unpubl.). Sampled populations of M. sativa (including M. capitata) from California are somewhat divergent in DNA sequences from sampled Chilean populations, in apparent conflict with earlier suggestions that M. sativa was recently introduced to North America from South America by Europeans (Baldwin, unpubl.). Madia sativa has been cultivated for seed-oil in South America, Europe, Africa, and Asia Minor (E. Zardini 1992). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Madia gracilis occurs widely in California (except the warm deserts), is scattered across much of Nevada, Oregon, and Washington (outside the driest regions), and extends into southernmost British Columbia, north-western Montana, and northern Utah. Near the coast, M. gracilis sometimes co-occurs with M. sativa; the two species are partially interfertile (M. gracilis tends to flower earlier than M. sativa; J. Clausen 1951). Reported occurrences of M. gracilis in Maine and South America have not been confirmed. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 21, p. 308. | FNA vol. 21, p. 308. |
Parent taxa | Asteraceae > tribe Heliantheae > subtribe Madiinae > Madia | Asteraceae > tribe Heliantheae > subtribe Madiinae > Madia |
Sibling taxa | ||
Synonyms | M. capitata | Sclerocarpus gracilis, M. gracilis subsp. collina, M. gracilis subsp. pilosa |
Name authority | Molina: Sag. Stor. Nat. Chili, 136. (1782) | (Smith) D. D. Keck: Madroño 5: 169. (1940) |
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