Madia sativa |
Madia elegans |
|
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Chile tarplant, Chile tarweed, Chilean tarplant, Chilean tarweed, coast tarweed, coastal tarweed |
autumn showy tarweed, common madia, showy tarweed |
|
Habit | Plants (0.3–)35–100(–240) cm, self-compatible (heads not showy). | Plants 6–250 cm, self-incompatible (heads showy). |
Stems | hirsute and glandular-pubescent, glands yellowish, purple, or black, lateral branches rarely surpassing main stems. |
proximally villous to hirsute, distally glandular-pubescent, glands yellowish, purple, or black, lateral branches sometimes surpassing main stems. |
Leaf | blades broadly lanceolate to linear-oblong or linear, 2–18 cm × 3–18(–29) mm. |
blades lanceolate to linear, 3–20 cm × 2–20 mm. |
Involucres | ovoid to urceolate, 6–16 mm. |
± globose to campanulate, 4.5–12 mm. |
Ray florets | (5–)8–13; corollas greenish yellow or sometimes purplish red abaxially or throughout, laminae 1.5–4 mm. |
(2–)5–22; corollas bright yellow (sometimes with maroon bases), laminae 4–20 mm. |
Disc florets | 11–14, bisexual, fertile; corollas 2–5 mm, pubescent; anthers ± dark purple. |
25–80+, functionally staminate; corollas 2.5–5 mm, pubescent; anthers yellow to brownish or ± dark purple. |
Phyllaries | hirsute and glandular-pubescent, glands yellowish, purple, or black, apices erect or ± reflexed, flat. |
± hirsute or villous, usually glandular-pubescent as well, glands yellowish, purple, or black, apices erect or reflexed, flat. |
Heads | in usually crowded, paniculiform, racemiform, or spiciform arrays. |
in open, corymbiform arrays. |
Disc cypselae | similar. |
0. |
Ray cypselae | black or brown, sometimes mottled, dull, compressed, beakless. |
black or brown, sometimes mottled, dull, compressed or ± 3-angled (slightly rounded abaxially, angled 15–45° adaxially), beakless (or nearly so). |
Paleae | mostly persistent, connate 1/2+ their lengths. |
mostly persistent, mostly connate 1/2+ their lengths. |
2n | = 32. |
= 16. |
Madia sativa |
Madia elegans |
|
Phenology | Flowering May–Oct. | Flowering Apr–Nov. |
Habitat | Grasslands, openings in shrublands and woods, disturbed sites, stream banks, roadsides | Grasslands, meadows, open sites in shrublands, woodlands, and forests, disturbed sites, often in coarse or clayey soils, sometimes serpentine |
Elevation | 0–1000 m (0–3300 ft) | 0–3400 m (0–11200 ft) |
Distribution |
CA; OR; WA; BC; South America (Argentina, Chile) [Pacific Islands (Hawaii, probably introduced)]
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CA; NV; OR; WA; Mexico (Baja California)
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Discussion | In North America, Madia sativa occurs on the Pacific Coast from California to British Columbia, sporadically in coastal ranges, and rarely eastward. Reports of M. sativa from Ontario and Quebec and from Alaska, Connecticut, Georgia, Idaho, Indiana, Iowa, Maine, Maryland, Massachusetts, New York, North Carolina, Pennsylvania, Vermont, and Wisconsin are putative waifs or misidentified M. glomerata. Molecular data and greenhouse studies have indicated that plants referable to M. capitata and M. sativa in California are not distinct (B. G. Baldwin, unpubl.). Sampled populations of M. sativa (including M. capitata) from California are somewhat divergent in DNA sequences from sampled Chilean populations, in apparent conflict with earlier suggestions that M. sativa was recently introduced to North America from South America by Europeans (Baldwin, unpubl.). Madia sativa has been cultivated for seed-oil in South America, Europe, Africa, and Asia Minor (E. Zardini 1992). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Madia elegans occurs widely in California outside the deserts and in southwestern Oregon and locally in western Nevada and Washington. It is unusually variable in morphology, ecology, and phenology. Molecular data have indicated that D. D. Keck’s (1959) infraspecific taxonomy for M. elegans needs revision. Putative natural (sterile) hybrids with M. sativa have been collected (e.g., D. D. Keck 2647, UC, from northern California). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 21, p. 308. | FNA vol. 21, p. 307. |
Parent taxa | Asteraceae > tribe Heliantheae > subtribe Madiinae > Madia | Asteraceae > tribe Heliantheae > subtribe Madiinae > Madia |
Sibling taxa | ||
Synonyms | M. capitata | M. elegans subsp. densifolia, M. elegans subsp. vernalis, M. elegans subsp. wheeleri |
Name authority | Molina: Sag. Stor. Nat. Chili, 136. (1782) | D. Don ex Lindley: Edwards’s Bot. Reg. 17: plate 1458. (1831) |
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