Madia sativa |
Madia citrigracilis |
|
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Chile tarplant, Chile tarweed, Chilean tarplant, Chilean tarweed, coast tarweed, coastal tarweed |
Shasta tarweed |
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Habit | Plants (0.3–)35–100(–240) cm, self-compatible (heads not showy). | Plants (10–)25–60 cm, self-compatible (heads not showy). |
Stems | hirsute and glandular-pubescent, glands yellowish, purple, or black, lateral branches rarely surpassing main stems. |
hirsute to villous, distally glandular-pubescent, glands yellowish, purple, or black, lateral branches often surpassing main stems (in large plants). |
Leaf | blades broadly lanceolate to linear-oblong or linear, 2–18 cm × 3–18(–29) mm. |
blades lanceolate to linear-oblong or linear, 3–15 cm × 2–14 mm. |
Involucres | ovoid to urceolate, 6–16 mm. |
± globose to ovoid or obovoid, 6–8 mm. |
Ray florets | (5–)8–13; corollas greenish yellow or sometimes purplish red abaxially or throughout, laminae 1.5–4 mm. |
5–8(–14); corollas pale yellow or greenish yellow, laminae 6–8 mm. |
Disc florets | 11–14, bisexual, fertile; corollas 2–5 mm, pubescent; anthers ± dark purple. |
3–10(–30), bisexual, fertile; corollas 2.5–3.5 mm, pubescent; anthers ± dark purple. |
Phyllaries | hirsute and glandular-pubescent, glands yellowish, purple, or black, apices erect or ± reflexed, flat. |
hirsute and glandular-pubescent, glands yellowish, purple, or black, apices erect or ± reflexed, flat. |
Heads | in usually crowded, paniculiform, racemiform, or spiciform arrays. |
in open, corymbiform arrays. |
Disc cypselae | similar. |
similar. |
Ray cypselae | black or brown, sometimes mottled, dull, compressed, beakless. |
black or brown, sometimes mottled, dull, compressed, beakless (or nearly so). |
Paleae | mostly persistent, connate 1/2+ their lengths. |
mostly persistent, mostly connate 1/2+ their lengths. |
2n | = 32. |
= 48. |
Madia sativa |
Madia citrigracilis |
|
Phenology | Flowering May–Oct. | Flowering Jun–Aug. |
Habitat | Grasslands, openings in shrublands and woods, disturbed sites, stream banks, roadsides | Openings in shrublands, woodlands, and forests |
Elevation | 0–1000 m (0–3300 ft) | 1400–2700 m (4600–8900 ft) |
Distribution |
CA; OR; WA; BC; South America (Argentina, Chile) [Pacific Islands (Hawaii, probably introduced)]
|
CA; NV |
Discussion | In North America, Madia sativa occurs on the Pacific Coast from California to British Columbia, sporadically in coastal ranges, and rarely eastward. Reports of M. sativa from Ontario and Quebec and from Alaska, Connecticut, Georgia, Idaho, Indiana, Iowa, Maine, Maryland, Massachusetts, New York, North Carolina, Pennsylvania, Vermont, and Wisconsin are putative waifs or misidentified M. glomerata. Molecular data and greenhouse studies have indicated that plants referable to M. capitata and M. sativa in California are not distinct (B. G. Baldwin, unpubl.). Sampled populations of M. sativa (including M. capitata) from California are somewhat divergent in DNA sequences from sampled Chilean populations, in apparent conflict with earlier suggestions that M. sativa was recently introduced to North America from South America by Europeans (Baldwin, unpubl.). Madia sativa has been cultivated for seed-oil in South America, Europe, Africa, and Asia Minor (E. Zardini 1992). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Madia citrigracilis occurs in northeastern California and northern Nevada. Small or young plants can be exceedingly difficult to distinguish morphologically from M. gracilis, which is one of two putative parental species suggested by J. Clausen et al. (1945) to have been involved in the hybrid (allopolyploid) origin of M. citrigracilis. Molecular data reinforce Clausen et al.’s hypothesis that M. citrigracilis is evolutionarily distinct from M. gracilis. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 21, p. 308. | FNA vol. 21, p. 307. |
Parent taxa | Asteraceae > tribe Heliantheae > subtribe Madiinae > Madia | Asteraceae > tribe Heliantheae > subtribe Madiinae > Madia |
Sibling taxa | ||
Synonyms | M. capitata | |
Name authority | Molina: Sag. Stor. Nat. Chili, 136. (1782) | D. D. Keck: Publ. Carnegie Inst. Wash. 564: 44. (1945) |
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