Macroptilium gibbosifolium |
Macroptilium |
|||||||||||||
---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
variableleaf bushbean |
bushbean |
|||||||||||||
Habit | Herbs, perennial; root turnip-shaped, small. | Herbs or vines, biennial or perennial, unarmed; roots turnip-shaped or elongate. | ||||||||||||
Stems | prostrate, solid, strigulose or hirsute. |
twining, erect, semierect, or prostrate, slender, sometimes hollow, sparsely to densely pubescent. |
||||||||||||
Leaves | stipules spreading, widely deltate or ovate, 4.5–8 × 2–2.7 mm; petiole 1.7–6 cm, strigose or hirsute; rachis 5–9 mm; stipels subulate, 1–5.5 mm; leaflets papery, blades orbiculate, ovate to broadly ovate or obovate, base attenuate or cuneate, apex acute or obtuse, surfaces strigose abaxially, hirtellous adaxially; lateral blades 2–4.3 × 1.6–2.9 cm, sometimes 2-lobed, terminal blades 2.3–4.7 × 1.6–2.8 cm. |
alternate, usually odd-pinnate, rarely unifoliolate; stipules present, persistent, not prolonged proximally, veined; petiolate; leaflets 3, stipels persistent, often reflexed, blade margins entire or 2-lobed, surfaces pubescent or glabrescent. |
||||||||||||
Inflorescences | 14–25 cm (cleistogamous inflorescence over soil, 1.5–2.2 mm, flowers 3.5 mm, corolla whitish), axes adpressed-pilose; tufts of bracts at base of peduncle mostly caducous; extrafloral nectaries inconspicuous at nodes; bracts and bracteoles subulate or lanceolate. |
6–12-flowered, axillary, pseudoracemes, cleistogamous flowers present in some species, with modified inflorescences, tufts of bracts present at base of peduncle, persistent or caducous, nodes swollen, glandular; node bracts present, caducous or inconspicuous, bracteoles present, usually caducous. |
||||||||||||
Pedicels | 0.7–1 mm. |
mostly shorter than calyx tube. |
||||||||||||
Flowers | 1 cm; calyx campanulate, 3.5–4.5 mm, hirtellous, calyx lobes as long as tube or longer; corolla salmon-orange; banner orbiculate, 5.5–7.5 × 4.5–6.5 mm; wings oblong-ovate, 5.1–10 × 3.3–6.6 mm; keel 4–8 mm; ovary nectary disc distally even; ovules 6. |
papilionaceous; calyx tubular or campanulate, lobes 5, lobes shorter to longer than tube, distinct or adaxial lobes partly connate; petals connate; corolla salmon-orange, red, or purple-black; banner orbiculate, obovate, or oblong, emarginate; wings ovate or oblong to obovate, conspicuously projected beyond distal bend of keel one directed upward to adopt function of banner; keel incurved, beak hooked, tip of beak hidden by wings; stamens 10, diadelphous; anthers sub-basifixed; ovary with nectary disc at base, style apically recurved and thickened, distal portion introrsely bearded, stigma terminal; ovules 4–24. |
||||||||||||
Fruits | legumes, patent or pendent, substipitate, valves plane or twisting, linear or oblong-falcate, dehiscent, non-septate, pilose, strigose, or hirsute. |
|||||||||||||
Legumes | pendent, oblong-falcate, 1.6–2.5 × 2.7–3 cm, valves not twisting at dehiscence. |
|||||||||||||
Seeds | 4 or 5, oblong, 2.9–3 × 2–2.2 mm. |
2–20, oblong or reniform; hilum not centric, oblong or ovate. |
||||||||||||
x | = 11. |
|||||||||||||
Macroptilium gibbosifolium |
Macroptilium |
|||||||||||||
Phenology | Flowering Jul–Sep. | |||||||||||||
Habitat | Rocky summits, oak forests, savanna-grasslands. | |||||||||||||
Elevation | 1300–2500 m. (4300–8200 ft.) | |||||||||||||
Distribution |
AZ; NM; TX; Mexico; Central America
|
Mexico; Central America; South America; s United States; West Indies |
||||||||||||
Discussion | Phaseolus micranthus M. Martens & Galeotti and P. parviflorus Schlechtendal are illegitimate names that pertain here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species 18 (4 in the flora). Flowers of Macroptilium species often become purple when dry; in some species, the banner is mostly pale green, with two basal auricles, wings are larger than banner and long-stipitate, brightly colored, and glabrous or pubescent; keel petals are distally connate and hooklike and basally adnate to the staminal tube. Vexillary stamens are distinct and often dilated at base. The ovary is subsessile, pubescent, and surrounded at the base by a cylindric, nectariferous disc; the style has an introrse beard and is apically recurved and thickened, with a terminal stigma. The greatest diversity in Macroptilium is found in South America (about 16 species; S. M. Espert et al. 2007); some species have been introduced for fodder and naturalized in the tropics and subtropics nearly worldwide. Molecular evidence establishes the monophylly of Macroptilium among the other New World genera of Phaseolinae (A. Delgado-Salinas et al. 2011). A diagnostic feature is the flower architecture: one wing petal is directed upward to adopt the function of the banner petal, which becomes a supporting structure of the wing petal and encloses the distally-hooked keel characteristic of all of the species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
||||||||||||
Key |
|
|||||||||||||
Source | FNA vol. 11. | FNA vol. 11. | ||||||||||||
Parent taxa | Fabaceae > subfam. Faboideae > Macroptilium | Fabaceae > subfam. Faboideae | ||||||||||||
Sibling taxa | ||||||||||||||
Subordinate taxa | ||||||||||||||
Synonyms | Phaseolus gibbosifolius, M. heterophyllum, M. heterophyllum var. rotundifolium, P. bilobatus, P. heterophyllus, P. heterophyllus var. rotundifolius, P. macropoides, P. macropus, P. rotundifolius, P. seleri | Phaseolus section macroptilium | ||||||||||||
Name authority | (Ortega) A. Delgado: Syst. Bot. 6: 295. (1981) | (Bentham) Urban: Symb. Antill. 9: 457. (1928) | ||||||||||||
Web links |