Macroptilium |
Macroptilium atropurpureum |
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bushbean |
conchito, purple bush-bean |
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Habit | Herbs or vines, biennial or perennial, unarmed; roots turnip-shaped or elongate. | |||||||||||||
Stems | twining, erect, semierect, or prostrate, slender, sometimes hollow, sparsely to densely pubescent. |
twining, solid, sericeous. |
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Leaves | alternate, usually odd-pinnate, rarely unifoliolate; stipules present, persistent, not prolonged proximally, veined; petiolate; leaflets 3, stipels persistent, often reflexed, blade margins entire or 2-lobed, surfaces pubescent or glabrescent. |
stipules retrorse, lanceolate or ovate, 5–6 × 1.8–2 mm; petiole 3.5–8 cm, canescent; rachis 5–8 mm; stipels linear, 1.2–2.5 mm; leaflets papery, blades elliptic or ovate, base cuneate or obtuse, apex acute, surfaces often canescent abaxially, strigose adaxially; lateral blades 3.3–4 × 2.7–3.3 cm, 2-lobed, terminal blades often hastate, 4.5–4.7 × 3–4.3 cm. |
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Inflorescences | 6–12-flowered, axillary, pseudoracemes, cleistogamous flowers present in some species, with modified inflorescences, tufts of bracts present at base of peduncle, persistent or caducous, nodes swollen, glandular; node bracts present, caducous or inconspicuous, bracteoles present, usually caducous. |
26–36 cm, axes pilose; tufts of bracts at base of peduncle persistent, 5.5 mm, lanceolate, canescent; extrafloral nectaries conspicuous at nodes; bracts and bracteoles lanceolate or subulate. |
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Pedicels | mostly shorter than calyx tube. |
1–2 mm. |
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Flowers | papilionaceous; calyx tubular or campanulate, lobes 5, lobes shorter to longer than tube, distinct or adaxial lobes partly connate; petals connate; corolla salmon-orange, red, or purple-black; banner orbiculate, obovate, or oblong, emarginate; wings ovate or oblong to obovate, conspicuously projected beyond distal bend of keel one directed upward to adopt function of banner; keel incurved, beak hooked, tip of beak hidden by wings; stamens 10, diadelphous; anthers sub-basifixed; ovary with nectary disc at base, style apically recurved and thickened, distal portion introrsely bearded, stigma terminal; ovules 4–24. |
1.7–3 cm; calyx campanulate, 6.2–8 mm, canescent, calyx lobes shorter than tube; corolla purple-black; banner oblong, 13–17 × 9–10.4 mm; wings obovate, 13–24 × 8–14.3 mm; keel 9–16 mm; ovary nectary disc distally oblique; ovules 13. |
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Fruits | legumes, patent or pendent, substipitate, valves plane or twisting, linear or oblong-falcate, dehiscent, non-septate, pilose, strigose, or hirsute. |
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Legumes | patent, linear, 7–10 × 0.3–0.4 cm, valves twisting at dehiscence. |
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Seeds | 2–20, oblong or reniform; hilum not centric, oblong or ovate. |
12 or 13, oblong, 3.5–5.2 × 2–3.2 mm. |
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Vines | , perennial; root elongate. |
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x | = 11. |
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2n | = 22. |
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Macroptilium |
Macroptilium atropurpureum |
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Phenology | Flowering Mar–Sep. | |||||||||||||
Habitat | introduced in tropical areas nearly worldwide.. | |||||||||||||
Elevation | 10–1200 m. (0–3900 ft.) | |||||||||||||
Distribution |
Mexico; Central America; South America; s United States; West Indies |
AZ; CA; FL; MS; NM; TX; Mexico; Central America; West Indies; South America (n Argentina); introduced in tropical areas nearly worldwide
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Discussion | Species 18 (4 in the flora). Flowers of Macroptilium species often become purple when dry; in some species, the banner is mostly pale green, with two basal auricles, wings are larger than banner and long-stipitate, brightly colored, and glabrous or pubescent; keel petals are distally connate and hooklike and basally adnate to the staminal tube. Vexillary stamens are distinct and often dilated at base. The ovary is subsessile, pubescent, and surrounded at the base by a cylindric, nectariferous disc; the style has an introrse beard and is apically recurved and thickened, with a terminal stigma. The greatest diversity in Macroptilium is found in South America (about 16 species; S. M. Espert et al. 2007); some species have been introduced for fodder and naturalized in the tropics and subtropics nearly worldwide. Molecular evidence establishes the monophylly of Macroptilium among the other New World genera of Phaseolinae (A. Delgado-Salinas et al. 2011). A diagnostic feature is the flower architecture: one wing petal is directed upward to adopt the function of the banner petal, which becomes a supporting structure of the wing petal and encloses the distally-hooked keel characteristic of all of the species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 11. | FNA vol. 11. | ||||||||||||
Parent taxa | ||||||||||||||
Sibling taxa | ||||||||||||||
Subordinate taxa | ||||||||||||||
Synonyms | Phaseolus section macroptilium | Phaseolus atropurpureus, P. atropurpureus var. canescens, P. atropurpureus var. pseuderythroloma, P. atropurpureus var. sericeus, P. canescens, P. dysophyllus, P. schiedeanus | ||||||||||||
Name authority | (Bentham) Urban: Symb. Antill. 9: 457. (1928) | (de Candolle) Urban: Symb. Antill. 9: 457. (1928) | ||||||||||||
Web links |