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mabrya

Habit Herbs, perennial; caudex woody.
Stems

pendent [erect], glandular-hairy.

Leaves

cauline, alternate;

petiole present;

blade fleshy, not leathery, margins broadly dentate [crenate].

Inflorescences

axillary, flowers solitary;

bracts absent.

Pedicels

present;

bracteoles absent.

Flowers

bisexual;

sepals 5, basally connate, calyx radially symmetric, campanulate, lobes ovate;

corolla ochroleucous to yellow [pink, red], bilaterally symmetric, bilabiate, tubular, tube base not spurred or gibbous, lobes 5, abaxial 3, adaxial 2;

stamens 4, basally adnate to corolla, didynamous, filaments glandular-hairy;

staminode 1, filamentous;

ovary 2-locular, placentation axile;

stigma 2-lobed.

Fruits

capsules, dehiscence loculicidal.

Seeds

20–300, dark brown [tan], oblong [ellipsoid, globular], wings absent [present].

x

= 12.

Mabrya

Distribution
from FNA
AZ; n Mexico
[BONAP county map]
Discussion

Species 5 (1 in the flora).

Mabrya is defined by several vegetative characteristics including pendent or erect, brittle stems, straight petioles, and glandular-hairy stems, leaves, and flowers. Species of Mabrya were placed previously in Maurandya Ortega by most authors; they have been recognized in Mabrya by W. J. Elisens (1985), D. A. Sutton (1988), and E. Fischer (2004). The phylogenetic relationships of Mabrya based on morphological characters are either unresolved (M. Ghebrehiwet et al. 2000) or poorly resolved (Elisens) because of character polymorphism and lack of synapomorphies for the genus. Molecular ITS data placed Holmgrenanthe in the Cymbalaria clade (M. Fernández-Mazuecos et al. 2013), in a subclade of seven genera that was sister to Asarina and Cymbalaria; Mabrya was sister to a node shared by Lophospermum and Maurandya.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 33. Author: Wayne J. Elisens.
Parent taxa Plantaginaceae
Subordinate taxa
M. acerifolia
Name authority Elisens: Syst. Bot. Monogr. 5: 57, fig. 26. (1985)
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