Lyonia |
Ericaceae subfam. vaccinioideae |
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maleberry, staggerbush |
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Habit | Shrubs or trees, (sometimes with woody burl, resprouting after fire). | |||||||||||||||||
Stems | erect to arching; twigs hairy, sometimes glabrescent (sometimes also lepidote). |
erect, arching, spreading, creeping, or procumbent. |
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Leaves | deciduous or persistent; blade elliptic, obovate, or ovate, membranous to coriaceous, margins entire, undulate, or serrulate [irregularly serrate], plane or revolute, surfaces multicellular, peltate-scaled or short-stalked stipitate-glandular-hairy, often unicellular-hairy on major veins or abaxial surface; venation brochidodromous (or reticulodromous). |
deciduous or persistent, alternate, sometimes pseudoverticillate (Pieris); petiole usually present, sometimes absent (some species of Vaccinium); blade plane, abaxial groove absent. |
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Inflorescences | axillary fascicles, panicles, or racemes, (2–)5–8(–12)-flowered, sometimes flowers solitary, (produced just before flowering). |
usually axillary, sometimes terminal, usually panicles or racemes, sometimes corymbs or fascicles, sometimes solitary flowers, (borne on leafy twigs, except Zenobia on leafless twigs); perulae absent; bracts much shorter than sepals (sometimes absent). |
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Pedicels | bracteoles 2, at or near base. |
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Flowers | sepals usually [4–]5[–8], slightly connate, ovate-deltate to lanceolate; petals usually [4–]5[–8], connate nearly their entire lengths, white to red, corolla cylindric to urceolate, lobes much shorter than tube, (sparsely stipitate-glandular-hairy or peltate-scaled); stamens usually [8–]10[–16], included; filaments geniculate, flattened, roughened or hairy, with or without spurs; anthers without awns, dehiscent by elliptic pores; pistil usually [4–]5[–8]-carpellate; ovary [4–]5[–8]-locular; (style slightly longer than stamens); stigma capitate-truncate. |
pendulous; perianth and androecium hypogynous or epigynous (Gaylussacia, Vaccinium); sepals (4-)5[-8]; petals 4-5(-6), connate (rarely distinct or nearly so in some species of Vaccinium), corolla deciduous, campanulate, cylindric, or urceolate, lobes usually much shorter (sometimes longer) than tube; intrastaminal nectary disc absent or present; stamens 8-10[-16]; anthers dehiscent by terminal pores or short slits; ovary 5- or 10-locular; placentation axile; style straight. |
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Fruits | capsular, globose to ovoid or ellipsoid, dry, (with [4–]5[–8] pale, decidedly thickened, whitish sutures). |
capsular, dehiscence loculicidal, or baccate or drupaceous, indehiscent. |
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Seeds | ca. 100–300, ellipsoidal, narrowly oblong, obovoid to angular-obovoid or narrowly conic, (sometimes tailed); testa cells elongate. |
2-300, distinct, ovoid or obovoid to ellipsoid, lanceoloid, or conic, to angular or wedge- or crescent-shaped, usually not winged, sometimes slightly winged or tailed. |
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x | = 12. |
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Lyonia |
Ericaceae subfam. vaccinioideae |
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Distribution |
e United States; Mexico; West Indies; e Asia |
Nearly worldwide; especially arctic; temperate; and alpine areas; also very diverse in neotropical cloud forests |
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Discussion | Arsenococcus Small; Desmothamnus Small; Neopieris Britton Species 36 (5 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera 46, species ca. 1600 (12 genera, 58 species in the flora) (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 8, p. 500. | FNA vol. 8, p. 496. | ||||||||||||||||
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Name authority | Nuttall: Gen. N. Amer. Pl. 1: 266. 1818, name conserved , | Arnott: M. Napier, Encycl. Brit. ed. 7 5: 118. (1832) — (as Vaccinieae) | ||||||||||||||||
Web links |