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common club-moss, elk-moss, lycopode à massue, running club-moss, running-pine, stag's horn clubmoss

club-moss, running-pine

Habit Plants mainly trailing on ground.
Roots

emerging from point of origin on underside of main stems.

Horizontal stems

on substrate surface.

on substrate surface or subterranean, long-creeping.

Gemmiferous branchlets and gemmae

absent.

Upright shoots

clustered, 0.6–1.2 cm diam., dominant main shoot with 3–6 branches mostly in lower 1/2.

scattered along horizontal stem, 5–16 mm diam., round or flat in cross section, unbranched or with 1–4 lateral branchlets.

Lateral branchlets

few and like upright shoots;

annual bud constrictions abrupt, branchlets mostly spreading.

Leaves

spreading, often somewhat ascending in distal 1/3 of branches, medium green, linear, 4–6 × 0.4–0.8 mm;

margins entire;

apex with narrow hair tip 2.5–4 mm.

not imbricate, linear to linear-lanceolate;

leaves on horizontal stems scattered, appressed, membranous;

leaves on lateral branchlets mostly 6-ranked or more, monomorphic with few exceptions, appressed, ascending to spreading, margins entire to dentate.

Peduncles

3.5–12.5 cm, with remote pseudowhorls of appressed leaves, loosely branched into 2–5 alternate stalks, 0.5–0.8 cm.

Strobili

2–5 on alternate stalks (if double, usually with stalks 5–8 mm), 15–25 × 3–6 mm.

single and sessile or multiple and pedunculate, apex blunt to acute;

peduncle, when present, conspicuously leafy;

sporophylls extremely reduced, much shorter than peduncle or stem leaves.

Sporangia

reniform.

Spores

reticulate, sides at equator convex, angles acute.

Sporophylls

1.5–2.5 mm, apex abruptly reduced to hair tip.

Gametophytes

nonphotosynthetic, mycorrhizal, subterranean, flat and irregularly button-shaped, with ring meristem around circumference.

x

= 34.

2n

= 68.

Lycopodium clavatum

Lycopodium

Habitat Fields and woods
Elevation 100–1800 m (300–5900 ft)
Distribution
from FNA
AK; CA; CT; GA; ID; IL; IN; KY; MA; MD; ME; MI; MN; MT; NC; NH; NJ; NY; OH; OR; PA; RI; TN; VA; VT; WA; WI; WV; BC; MB; NB; NF; NS; ON; PE; QC; SK; SPM; Mexico; Central America; South America; West Indies; Europe; Asia; Africa; Pacific Islands
[WildflowerSearch map]
[BONAP county map]
from USDA
Mainly temperate and subarctic
[BONAP county map]
Discussion

Plants found in eastern North America have been called Lycopodium clavatum var. clavatum; those in the western part of the range, which have been called L. clavatum var. integrifolium Goldie, are distinguished by early shedding of the characteristic hairs on the leaf tips.

Lycopodium dendroideum group

(Discussion copyrighted by Flora of North America; reprinted with permission.)

In striking contrast to Diphasiastrum, Huperzia, and Lycopodiella, interspecific hybridization is practically unknown in Lycopodium. Many of the species now recognized in Lycopodium have been segregated from Lycopodium clavatum, L. annotinum, and L. jussiaei Desvaux ex Poiret. The three groups given in the key below should probably be treated as subgenera.

Species 15–25 (6 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Strobili pedunculate; upright shoots with 2–5 branches, not treelike; leaves with hair tips 1–4 mm (these may fall off early, but remain at shoot apices) (L. clavatum group).
→ 2
1. Strobili sessile; upright shoots either unbranched or much branched to produce treelike habit; leaves lacking hair tips.
→ 3
2. Strobili mostly solitary on peduncle, if paired then nearly lacking pedicels; leaves 3–5 mm, ascending to appressed; branches 2–3(–4), mostly upright.
L. lagopus
2. Strobili 2–5, borne on loosely alternate pedicels, 0.5–0.8 cm; leaves 4–6 mm, spreading to somewhat ascending; branches 3–6, mostly oblique or spreading.
L. clavatum
3. Strobili single at top of upright shoot; shoot unbranched or branched 1–2 times; horizontal stems on substrate surface (L. annotinum group).
L. annotinum
3. Strobili 1–7 at top of many-branched, upright, treelike shoot; horizontal stems subterranean (L. dendroideum group).
→ 4
4. Lateral shoots flat in cross section, leaves unequal in size, lateral leaves spreading and twisted, adaxial surfaces facing upward, proximal leaves much reduced; leaves on main axis dark green, tightly appressed.
L. obscurum
4. Lateral shoots round in cross section, leaves equal in size, none twisted, adaxial leaf surfaces all facing stem, proximal leaves not reduced; leaves on main axis light or dark green, spreading or appressed.
→ 5
5. Leaf ranks 1 on upperside of lateral branch, 2 on each side, and 1 on underside; leaves of main axis below branches dark green, tightly appressed, soft to touch.
L. hickeyi
5. Leaf ranks 2 on top of lateral branch, 1 on each side, and 2 on underside; leaves of main axis below branches pale green, spreading, prickly to touch.
L. dendroideum
Source FNA vol. 2. FNA vol. 2.
Parent taxa Lycopodiaceae > Lycopodium Lycopodiaceae
Sibling taxa
L. annotinum, L. dendroideum, L. hickeyi, L. lagopus, L. obscurum
Subordinate taxa
L. annotinum, L. clavatum, L. dendroideum, L. hickeyi, L. lagopus, L. obscurum
Synonyms L. clavatum var. subremotum
Name authority Linnaeus: Sp. Pl. 2: 1101. (1753) Linnaeus: Sp. Pl. 2: 1100. 1753; Gen. Pl. ed. 5, 486, (1754)
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