Lycopodium clavatum |
Lycopodiaceae |
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common club-moss, elk-moss, lycopode à massue, running club-moss, running-pine, stag's horn clubmoss |
club-moss family |
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Habit | Plants terrestrial, on rock, or epiphytic. | |||||||||||||||||||||||||
Roots | emerging near origin, or growing through cortex and emergent some distance from origin. |
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Horizontal stems | on substrate surface. |
present or absent, mainly protostelic, in some species becoming actino- or plectostelic, on substrate surface or subterranean, or forming stolons. |
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Lateral shoots | present or absent, simple or branched, branching pattern dichotomous and sometimes pseudomonopodial; leaves uniform or dimorphic or trimorphic. |
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Upright shoots | clustered, 0.6–1.2 cm diam., dominant main shoot with 3–6 branches mostly in lower 1/2. |
simple or branched, usually conspicuously leafy at least at base; abscising gemmae formed by reduced lateral shoots. |
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Upright and lateral shoots | round or flat in cross section; leaves on subterranean parts flat, appressed, nonphotosynthetic, and scalelike; leaves on aerial parts appressed, ascending, or spreading, with 1 central unbranched vein, needlelike to lanceolate to ovate, remote to dense and imbricate, with or without basal and/or mucilage canals. |
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Lateral branchlets | few and like upright shoots; annual bud constrictions abrupt, branchlets mostly spreading. |
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Leaves | spreading, often somewhat ascending in distal 1/3 of branches, medium green, linear, 4–6 × 0.4–0.8 mm; margins entire; apex with narrow hair tip 2.5–4 mm. |
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Peduncles | 3.5–12.5 cm, with remote pseudowhorls of appressed leaves, loosely branched into 2–5 alternate stalks, 0.5–0.8 cm. |
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Strobili | 2–5 on alternate stalks (if double, usually with stalks 5–8 mm), 15–25 × 3–6 mm. |
sessile or stalked, upright, nodding, or pendent. |
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Sporangia | solitary, adaxial near leaf base or axillary; subtending leaves (sporophylls) unmodified and photosynthetic to much modified, nonphotosynthetic, reduced, and aggregated in strobili; sporangia reniform to globose, thick-walled with hundreds of spores, outer walls variously modified. |
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Spores | all 1 kind, trilete, thick-walled, surfaces pitted to small-grooved, rugulate, or reticulate. |
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Sporophylls | 1.5–2.5 mm, apex abruptly reduced to hair tip. |
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Gametophytes | subterranean and nonphotosynthetic or surficial and photosynthetic. |
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2n | = 68. |
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Lycopodium clavatum |
Lycopodiaceae |
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Habitat | Fields and woods | |||||||||||||||||||||||||
Elevation | 100–1800 m (300–5900 ft) | |||||||||||||||||||||||||
Distribution |
AK; CA; CT; GA; ID; IL; IN; KY; MA; MD; ME; MI; MN; MT; NC; NH; NJ; NY; OH; OR; PA; RI; TN; VA; VT; WA; WI; WV; BC; MB; NB; NF; NS; ON; PE; QC; SK; SPM; Mexico; Central America; South America; West Indies; Europe; Asia; Africa; Pacific Islands
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Worldwide |
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Discussion | Plants found in eastern North America have been called Lycopodium clavatum var. clavatum; those in the western part of the range, which have been called L. clavatum var. integrifolium Goldie, are distinguished by early shedding of the characteristic hairs on the leaf tips. Lycopodium dendroideum group (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The Lycopodiaceae are an extremely diverse, ancient family. The family may contain even more than the estimated 400 species because the tropical members and the very large genus Phlegmariurus are still poorly known. The relationships among genera of Lycopodiaceae are not well understood because large evolutionary gaps exist among most genera. Some of the genera, notably Diphasiastrum, Huperzia, and Lycopodiella, exhibit extensive interspecific hybridization, which has caused much taxonomic confusion in the past. Differences in expressions of many of the generic characters are subtle, and some of the characters are microscopic. Genera 10–15, species 350–400 (7 genera, 27 species in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 2. | FNA vol. 2, p. 18. | ||||||||||||||||||||||||
Parent taxa | Lycopodiaceae > Lycopodium | |||||||||||||||||||||||||
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Synonyms | L. clavatum var. subremotum | |||||||||||||||||||||||||
Name authority | Linnaeus: Sp. Pl. 2: 1101. (1753) | Mirbel | ||||||||||||||||||||||||
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