Lycopodium annotinum |
Lycopodium hickeyi |
|
---|---|---|
bristly club-moss, lycopode interrompu, stiff club-moss |
Hickey's tree club-moss, lycopode de Hickey, Pennsylvania clubmoss |
|
Horizontal stems | on substrate surface. |
subterranean. |
Upright shoots | clustered, mainly unbranched or sparsely branching mainly at base, 1.2–1.6 cm diam.; annual bud constrictions abrupt and conspicuous. |
treelike, many branched, branchlets numerous and strongly differentiated; annual bud constrictions absent; leaves on main axis below lateral branchlets tightly appressed, dark green, needlelike, 3.5–4.5 × 0.5–0.6 mm, soft. |
Lateral branchlets | few and like upright shoots but annual bud constrictions absent. |
round in cross section, 4–7 mm diam.; annual bud constrictions inconspicuous; leaves ascending, in 6 ranks, 1 on upperside, 4 lateral, and 1 on underside, equal in size, linear, widest in middle; margins entire; apex acuminate, lacking hair tip. |
Leaves | spreading to reflexed, dark green, linear-lanceolate, (2.5–)5–8 × 0.6–1.2 mm; margins closely and shallowly dentate mainly in distal 1/2; apex sharply pointed, lacking hair tip. |
|
Strobili | solitary, sessile on shoots, 15–30 × 3.5–4.5 mm. |
sessile, 1–7 per upright shoot, 15–65 mm. |
Sporophylls | (1.5–)3.5 × 0.7(–2) mm, abruptly narrowed to pointed tip. |
3–3.5 × 2–2.5 mm, apex long, gradually narrowing to tip. |
2n | = 68. |
= 68. |
Lycopodium annotinum |
Lycopodium hickeyi |
|
Habitat | Swampy or moist coniferous forests, mountain forests, and exposed grassy or rocky sites | Mainly in hardwood forests and second-growth, shrubby habitats |
Elevation | 0–1850 m (0–6100 ft) | 0–1600 m (0–5200 ft) |
Distribution |
AK; AZ; CO; CT; ID; KY; MA; MD; ME; MI; MN; MT; NC; NH; NJ; NM; NY; OH; OR; PA; RI; TN; UT; VA; VT; WA; WI; WY; AB; BC; MB; NB; NF; NS; NT; ON; PE; QC; SK; YT; SPM; Greenland |
CT; IN; KY; MA; MD; ME; MI; MN; NC; NH; NJ; NY; OH; PA; RI; TN; VA; VT; WI; WV; NB; NF; NS; ON; PE; QC; SK
|
Discussion | This widespread and common club-moss has been divided into various forms or varieties, some of which have been treated as species. Present evidence supports the hypothesis that these are environmentally induced forms, the most distinctive of which has been called Lycopodium annotinum var. alpestre C. Hartman, with leaves only 2.5–6 mm, very leathery, entire-margined, and appressed. Plants intermediate between this and L. annotinum var. annotinum are a form that has been called var. pungens (Bachelot de la Pylaie) Desvaux, an invalid name. Both are found in cold, bleak, northern or high elevation habitats. The species should be studied in detail to determine whether it contains any groups that should be recognized taxonomically. Lycopodium clavatum group (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The range of Lycopodium hickeyi overlaps with that of L. obscurum and extends considerably north and west of that species. Although the arrangement of the leaf ranks is similar to that of L. obscurum, the leaf dimorphy and the ascending orientation and absence of twisting of the leaves are diagnostic. Where ranges of two or three species overlap, indivual species retain their identities, indicating that their critical differences have a genetic basis. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Lycopodiaceae > Lycopodium | Lycopodiaceae > Lycopodium |
Sibling taxa | ||
Synonyms | L. obscurum var. isophyllum | |
Name authority | Linnaeus: Sp. Pl. 2: 1103. (1753) | W. H. Wagner: Beitel, & R. C. Moran, Amer. Fern J. 79: 119–121. (1989) |
Web links |