Lycopodium annotinum |
Lycopodiaceae |
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bristly club-moss, lycopode interrompu, stiff club-moss |
club-moss family |
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Habit | Plants terrestrial, on rock, or epiphytic. | |||||||||||||||||||||||||
Roots | emerging near origin, or growing through cortex and emergent some distance from origin. |
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Horizontal stems | on substrate surface. |
present or absent, mainly protostelic, in some species becoming actino- or plectostelic, on substrate surface or subterranean, or forming stolons. |
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Lateral shoots | present or absent, simple or branched, branching pattern dichotomous and sometimes pseudomonopodial; leaves uniform or dimorphic or trimorphic. |
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Upright shoots | clustered, mainly unbranched or sparsely branching mainly at base, 1.2–1.6 cm diam.; annual bud constrictions abrupt and conspicuous. |
simple or branched, usually conspicuously leafy at least at base; abscising gemmae formed by reduced lateral shoots. |
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Upright and lateral shoots | round or flat in cross section; leaves on subterranean parts flat, appressed, nonphotosynthetic, and scalelike; leaves on aerial parts appressed, ascending, or spreading, with 1 central unbranched vein, needlelike to lanceolate to ovate, remote to dense and imbricate, with or without basal and/or mucilage canals. |
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Lateral branchlets | few and like upright shoots but annual bud constrictions absent. |
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Leaves | spreading to reflexed, dark green, linear-lanceolate, (2.5–)5–8 × 0.6–1.2 mm; margins closely and shallowly dentate mainly in distal 1/2; apex sharply pointed, lacking hair tip. |
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Strobili | solitary, sessile on shoots, 15–30 × 3.5–4.5 mm. |
sessile or stalked, upright, nodding, or pendent. |
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Sporangia | solitary, adaxial near leaf base or axillary; subtending leaves (sporophylls) unmodified and photosynthetic to much modified, nonphotosynthetic, reduced, and aggregated in strobili; sporangia reniform to globose, thick-walled with hundreds of spores, outer walls variously modified. |
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Spores | all 1 kind, trilete, thick-walled, surfaces pitted to small-grooved, rugulate, or reticulate. |
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Sporophylls | (1.5–)3.5 × 0.7(–2) mm, abruptly narrowed to pointed tip. |
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Gametophytes | subterranean and nonphotosynthetic or surficial and photosynthetic. |
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2n | = 68. |
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Lycopodium annotinum |
Lycopodiaceae |
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Habitat | Swampy or moist coniferous forests, mountain forests, and exposed grassy or rocky sites | |||||||||||||||||||||||||
Elevation | 0–1850 m (0–6100 ft) | |||||||||||||||||||||||||
Distribution |
AK; AZ; CO; CT; ID; KY; MA; MD; ME; MI; MN; MT; NC; NH; NJ; NM; NY; OH; OR; PA; RI; TN; UT; VA; VT; WA; WI; WY; AB; BC; MB; NB; NF; NS; NT; ON; PE; QC; SK; YT; SPM; Greenland |
Worldwide |
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Discussion | This widespread and common club-moss has been divided into various forms or varieties, some of which have been treated as species. Present evidence supports the hypothesis that these are environmentally induced forms, the most distinctive of which has been called Lycopodium annotinum var. alpestre C. Hartman, with leaves only 2.5–6 mm, very leathery, entire-margined, and appressed. Plants intermediate between this and L. annotinum var. annotinum are a form that has been called var. pungens (Bachelot de la Pylaie) Desvaux, an invalid name. Both are found in cold, bleak, northern or high elevation habitats. The species should be studied in detail to determine whether it contains any groups that should be recognized taxonomically. Lycopodium clavatum group (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The Lycopodiaceae are an extremely diverse, ancient family. The family may contain even more than the estimated 400 species because the tropical members and the very large genus Phlegmariurus are still poorly known. The relationships among genera of Lycopodiaceae are not well understood because large evolutionary gaps exist among most genera. Some of the genera, notably Diphasiastrum, Huperzia, and Lycopodiella, exhibit extensive interspecific hybridization, which has caused much taxonomic confusion in the past. Differences in expressions of many of the generic characters are subtle, and some of the characters are microscopic. Genera 10–15, species 350–400 (7 genera, 27 species in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 2. | FNA vol. 2, p. 18. | ||||||||||||||||||||||||
Parent taxa | Lycopodiaceae > Lycopodium | |||||||||||||||||||||||||
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Name authority | Linnaeus: Sp. Pl. 2: 1103. (1753) | Mirbel | ||||||||||||||||||||||||
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