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Piper's wood-rush, smooth woodrush

hairy wood rush, wood rush

Habit Herbs, perennial, usually cespitose, often with short, mostly vertical to running rhizomes and/or (less commonly) stolons.
Rhizomes

horizontal, short.

Culms

densely cespitose, 10–30(–35) cm.

round.

Cataphylls

absent.

Leaves

basal blade green, 5–10 cm × 2–4 mm, firm, essentially glabrous;

cauline leaves 2–3, 3–7 cm × 3–5 mm.

sheaths closed, without auricles at throat (junction with blade), usually pilose;

blade flat or channeled, never septate, margins with long, soft, multicellular hairs, apex often thickened (callous), veins commonly indistinct.

Inflorescences

branches spreading less than 90°, lax;

proximal inflorescence bract leaflike, 0.8–1.5 cm;

bracts and bracteoles brown, clear at apex, margins strongly ciliate.

terminal;

flowers inserted individually or in dense clusters (glomerules) variously arranged;

bracts subtending inflorescence (proximal inflorescence bracts) 2, mostly leaflike;

bracts subtending inflorescence branches 1–2, reduced;

bracteoles subtending flowers 2–3.

Flowers

single or in clusters of 2–3;

tepals dark brown, 1–2.5 mm, ± equal, apex acute, not reflexed;

anthers ± equaling filament length;

stigmas 5 times style length.

tepals 6, in 2 whorls;

stamens 6.

Capsules

dark brown, ellipsoid, shorter than 2.5 mm, longer than tepals;

beak absent.

1-locular, generally globose;

beak often formed by persistent style base.

Seeds

light yellow-brown, lanceolate, narrowed at ends, 1.2 mm.

3, globose to ovoid, base often with tuft of fibrous hairs (vestige of funiculus);

nutritive appendage from outer seed coat (caruncle) often present, white, barely visible to ± equaling seed body.

x

= 6.

2n

= 24.

Luzula piperi

Luzula

Phenology Flowering and fruiting summer.
Habitat Snowbeds and mesic heaths in subalpine and oceanic zones
Elevation 400–2400 m (1300–7900 ft)
Distribution
from FNA
AK; ID; MT; OR; WA; AB; BC; YT; e Asia
[WildflowerSearch map]
[BONAP county map]
from USDA
Temperate and arctic regions worldwide; tropical mountains
[BONAP county map]
Discussion

The leaves of Luzula are primarily basal; cauline leaves are usually reduced.

Luzula species have diffuse centromeres and small chromosomes. That has resulted in much confusion in interpretation and reporting of chromosome counts. No attempt has been made to include reported counts that could not reasonably be verified by the author.

Excluded species: Luzula sudetica (Willdenow) de Candolle. Although reports of this European species appear frequently in the North American literature, I have seen no specimens that confirm its presence. No chromosome counts are published for North American material. Since this species has a distinct cytotype, 2n = 48 (H. Nordenskiöld 1956), it should not be difficult to verify on this basis.

Species ca. 108 (23 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Flowers in dense clusters (glomerules); inflorescences spikelike or umbellate; seeds with caruncle conspicuous to barely visible
subg. Luzula
1. Flowers solitary or in small clusters of 2–4; inflorescences mostly unbranched or dichasial; seeds with caruncle conspicuous to absent.
→ 2
2. Flowers solitary; inflorescences corymbose, rarely branching; seeds with caruncle conspicuous
subg. Pterodes
2. Flowers mostly in pairs, rarely in clusters of 3–4, or solitary; inflorescences paniculate or dichasial; seeds with caruncle inconspicuous or absent
subg. Anthelaea
Source FNA vol. 22. FNA vol. 22, p. 255. Author: Janice Coffey Swab.
Parent taxa Juncaceae > Luzula > subg. Anthelaea Juncaceae
Sibling taxa
L. acuminata, L. arctica, L. arcuata, L. bulbosa, L. campestris, L. comosa, L. confusa, L. divaricata, L. echinata, L. groenlandica, L. hitchcockii, L. kjellmaniana, L. luzuloides, L. multiflora, L. orestera, L. pallidula, L. parviflora, L. rufescens, L. spicata, L. subcapitata, L. subcongesta, L. wahlenbergii
Subordinate taxa
L. subg. Anthelaea, L. subg. Luzula, L. subg. Pterodes
Synonyms Juncoides piperi, L. wahlenbergii subsp. piperi Juncoides
Name authority (Coville) M. E. Jones: Bull. Biol. Ser. Bull. State Univ. Montana 15: 22. (1910) de Candolle: in J. Lamarck and A. P. de Candolle, Fl. France, ed. 3 1: 198; 3: 158. (1805)
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