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arctic wood rush

hairy wood rush, wood rush

Habit Herbs, perennial, usually cespitose, often with short, mostly vertical to running rhizomes and/or (less commonly) stolons.
Culms

densely cespitose, 5-20 cm.

round.

Cataphylls

absent.

Leaves

sheaths brown to straw-colored;

basal leaves to 10 cm × 4 mm;

cauline leaves usually 2, reduced.

sheaths closed, without auricles at throat (junction with blade), usually pilose;

blade flat or channeled, never septate, margins with long, soft, multicellular hairs, apex often thickened (callous), veins commonly indistinct.

Inflorescences

glomerules 1-3, sessile;

proximal inflorescence bract inconspicuous, brown, much shorter to ± equaling inflorescence, apex often clear, dentate;

bracts deep brown, margins dentate;

bracteoles deep brown, margins dentate.

terminal;

flowers inserted individually or in dense clusters (glomerules) variously arranged;

bracts subtending inflorescence (proximal inflorescence bracts) 2, mostly leaflike;

bracts subtending inflorescence branches 1–2, reduced;

bracteoles subtending flowers 2–3.

Flowers

tepals deep brown with narrow clear margins and apex, 1.7-2.1 mm;

anthers ± equaling filament length.

tepals 6, in 2 whorls;

stamens 6.

Capsules

dark reddish to blackish, shining, spheric, 1.8-2.1 mm, usually exceeding tepals.

1-locular, generally globose;

beak often formed by persistent style base.

Seeds

translucent, clear brown, broadly elliptic, with few entangled hairs, 1-1.2 mm.

3, globose to ovoid, base often with tuft of fibrous hairs (vestige of funiculus);

nutritive appendage from outer seed coat (caruncle) often present, white, barely visible to ± equaling seed body.

x

= 6.

2n

= 24.

Luzula arctica

Luzula

Phenology Flowering and fruiting summer.
Habitat Wet, stony places on slopes and in dwarf shrub heaths in alpine and arctic tundra; circumpolar.
Elevation 0-1200 m (0-3900 ft)
Distribution
from FNA
AK; BC; LB; MB; NT; NU; QC; YT; Greenland; Eurasia
from USDA
Temperate and arctic regions worldwide; tropical mountains
[BONAP county map]
Discussion

The leaves of Luzula are primarily basal; cauline leaves are usually reduced.

Luzula species have diffuse centromeres and small chromosomes. That has resulted in much confusion in interpretation and reporting of chromosome counts. No attempt has been made to include reported counts that could not reasonably be verified by the author.

Excluded species: Luzula sudetica (Willdenow) de Candolle. Although reports of this European species appear frequently in the North American literature, I have seen no specimens that confirm its presence. No chromosome counts are published for North American material. Since this species has a distinct cytotype, 2n = 48 (H. Nordenskiöld 1956), it should not be difficult to verify on this basis.

Species ca. 108 (23 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Flowers in dense clusters (glomerules); inflorescences spikelike or umbellate; seeds with caruncle conspicuous to barely visible
subg. Luzula
1. Flowers solitary or in small clusters of 2–4; inflorescences mostly unbranched or dichasial; seeds with caruncle conspicuous to absent.
→ 2
2. Flowers solitary; inflorescences corymbose, rarely branching; seeds with caruncle conspicuous
subg. Pterodes
2. Flowers mostly in pairs, rarely in clusters of 3–4, or solitary; inflorescences paniculate or dichasial; seeds with caruncle inconspicuous or absent
subg. Anthelaea
Source FNA vol. 22, p. 263. FNA vol. 22, p. 255. Author: Janice Coffey Swab.
Parent taxa Juncaceae > Luzula > subg. Luzula Juncaceae
Sibling taxa
L. acuminata, L. arcuata, L. bulbosa, L. campestris, L. comosa, L. confusa, L. divaricata, L. echinata, L. groenlandica, L. hitchcockii, L. kjellmaniana, L. luzuloides, L. multiflora, L. orestera, L. pallidula, L. parviflora, L. piperi, L. rufescens, L. spicata, L. subcapitata, L. subcongesta, L. wahlenbergii
Subordinate taxa
L. subg. Anthelaea, L. subg. Luzula, L. subg. Pterodes
Synonyms L. nivalis Juncoides
Name authority Blytt: in M. N. Blytt and A. G. Blytt, Norges Flora 1: 299. (1861) de Candolle: in J. Lamarck and A. P. de Candolle, Fl. France, ed. 3 1: 198; 3: 158. (1805)
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