Ludwigia spathulata |
Ludwigia alternifolia |
|
---|---|---|
southern water purslane, spoon primrose-willow |
bushy seedbox, ludwigie à feuilles alternes, rattlebox, seedbox, square-pod water-primrose |
|
Habit | Herbs creeping and rooting at nodes, often forming mats. | |
Roots | fusiform, fascicled, thickened, epidermis splitting or peeling near base. |
|
Stems | prostrate or decumbent and ascending distally, slightly ridged, well branched, 10–40cm, densely strigillose throughout. |
subterete or somewhat angled, with narrow raised lines or wings decurrent from leaf axils, 40–150 cm, well branched in distal 1/2, glabrous or sparsely to densely strigillose. |
Leaves | opposite; stipules narrowly deltate or ovate, 0.05–0.15 × 0.05–0.1 mm; petiole very narrowly winged, 0.3–0.9 cm, blade elliptic-spatulate or narrowly so, 0.9–1.7 × 0.3–0.9 cm, base attenuate, margins entire, apex acute, surfaces strigillose; bracts not reduced except at branch tips. |
stipules narrowly deltate, 0.05–0.2 × 0.05–0.1 mm; petiole 0.1–0.3(–0.7) cm; blade lanceolate-elliptic, (0.6–)2–12 × (0.3–)1–1.5(–2.5) cm, base attenuate, margins entire, apex acute, surfaces strigillose throughout or glabrate with strigillose veins; bracts often reduced and more linear. |
Inflorescences | leafy spikes or racemes, flowers usually paired in leaf axils; bracteoles attached at base of ovary or on short pedicel, narrowly oblong or oblanceolate, 0.2–0.8 × 0.05–0.2 mm, apex acute, often obscured by hairs. |
leafy racemes, flowers solitary in leaf axils; bracteoles linear-lanceolate, 1–2.5 mm, margins entire, apex acute or subacuminate, glabrous or with scattered hairs, attached near base of ovary. |
Flowers | sepals ascending, pale green, broadly ovate-deltate, 1–1.7 × 1.1–1.7 mm, margins entire, apex acuminate, surfaces densely strigillose; petals 0; filaments translucent, 0.5–0.8 mm, anthers 0.2–0.4 × 0.3–0.5 mm; pollen shed singly; ovary oblong-obovoid, 4-angled to subterete, 1.5–2.5 × 1–1.5 mm; nectary disc elevated 0.1–0.2 mm on ovary apex, yellowish green, 0.7–0.9 mm diam., 4-lobed, glabrous; style yellowish green, 0.3–0.5 mm, glabrous, stigma pale yellow, capitate, 0.2–0.3 × 0.2–0.3 mm, not exserted beyond anthers. |
sepals narrowly ovate-deltate, (6–)6.5–9.5 × 4–6.5 mm, margins entire, apex acute to obtuse, surfaces strigillose, sometimes mixed with villous hairs, or glabrate; petals cordate, 10–14 ×8–12 mm, base attenuate, apex emarginate; filaments opaque white, awl-shaped, 1–3 mm, anthers 1–1.7 × 0.6–0.8 mm; ovary subcuboid to globose, 2.5–4 ×2.5–3.5 mm; nectary disc slightly elevated on ovary apex, 0.8–1.5 mm diam., 4-lobed, ringed with soft, curly hairs or glabrous; style 1.5–2.4 mm, glabrous, stigma light yellow, capitate to hemispherical, 1–1.3 × 1.4–2 mm, shallowly 4-lobed, not or scarcely exserted beyond anthers. |
Capsules | oblong-obovoid, subterete, 2.5–4(–4.5) × 1.5–2.5 mm, thin-walled, seeds often visible on exocarp as small bumps, irregularly dehiscent or dispersing as unit, pedicel 0–0.5 mm. |
subcuboid to squarish globose, 4–6(–7) × 4–6 mm, 4-angled and 4-winged, wings 0.3–1.5 mm wide, pedicel 2–7 mm. |
Seeds | dark reddish brown, ellipsoid, 0.5–0.7 × 0.4–0.5 mm, surface cells transversely elongate. |
light brown, oblong to reniform, 0.5–0.8 × 0.2–0.4 mm, surface cells elongate transversely to seed length. |
2n | = 32. |
= 16. |
Ludwigia spathulata |
Ludwigia alternifolia |
|
Phenology | Flowering summer–early fall. | |
Habitat | Swamps, damp, peaty places, roadside ditches, margins of cultivated fields. | |
Elevation | 0–1800 m. (0–5900 ft.) | |
Distribution |
AL; AR; CO; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; MI; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; VT; WI; WV; ON; QC
|
|
Discussion | Flowers May–Sep. Ditches, swales, edges of ponds, lakes, sinks, swamps, sandy river bars, dried seasonal ponds, disturbed low savannas; of conservation concern; 0–200 m; Ala., Fla., Ga., S.C. The tetraploid Ludwigia spathulata is relatively uncommon and occurs primarily on the Gulf Coastal Plain in the panhandle of Florida, southern Alabama, and southwestern Georgia. Outlying populations have also been collected in transitional areas between the Coastal Plain and the Piedmont in South Carolina and Georgia. With its small apetalous flowers, L. spathulata is modally autogamous and shows low morphological variability. Its strongest affinities appear to be with L. palustris, with which it shares a genome (C. I. Peng 1988, 1989). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Ludwigia alternifolia is widespread and common in the eastern half of the flora area, as far west as Ontario in Canada, eastern Colorado, and Texas. Ludwigia macrocarpa Michaux 1803 is a superfluous, illegitimate name for L. alternifolia. Rhexia linearifolia Poiret was originally described as a species of Melastomataceae, but the description is superficially similar to Ludwigia and has often been included in synonymy with L. alternifolia. The type of R. linearifolia has not been located or studied. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 10. | FNA vol. 10. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Isnardia spathulata | Isnardia alternifolia, I. alternifolia var. salicifolia, I. alternifolia var. uniflora, I. aurantiaca, L. alternifolia var. linearifolia, L. alternifolia var. pubescens, L. angustifolia var. ramosissima, L. aurantiaca, L. ramosissima, L. salicifolia, L. uniflora |
Name authority | Torrey & A. Gray: Fl. N. Amer. 1: 526. (1840) | Linnaeus: Sp. Pl. 1: 118. (1753) |
Web links |