The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

hairy primrose-willow

hairy seedbox, Rafinesque's seedbox, spindleroot

Habit Herbs often with prominent aerenchyma when base submerged, forming stolons 30–250 cm, 2–4 mm thick, creeping in mud or floating in water, sometimes bearing flowers and fruits.
Roots

fusiform, somewhat thickened, often fascicled.

Stems

erect, subterete, densely branched, 40–120 cm, densely hirtellous.

subterete to angled, with raised lines or narrow wings decurrent from leaf axils, (30–)50–120 cm, well branched in distal 1/2, densely erect-hirsute to sometimes glabrous.

Leaves

alternate;

stipules ovate to lanceolate, 0.2–0.25 × 0.1–0.15 mm, usually obscured by pubescence;

stolons: petiole 0.2–0.7 cm, blade obovate or elliptic to orbiculate, 0.6–2 × 0.5–1.1 cm, margins with distinct hydathodal teeth, base attenuate, surfaces densely hirtellous to glabrate;

stems: petiole 0–0.2(–1) cm, blade elliptic or lanceolate-elliptic to very narrowly elliptic, 1.5–8(–10) × 0.3–1.2(–1.4) cm, base cuneate or attenuate, margins entire with obscure hydathodal glands, apex acute or narrowly acute, surfaces ± densely hirtellous, leaves on branches much reduced;

bracts much reduced.

stipules narrowly deltate, 0.05–0.15 × 0.05–0.1 mm;

sessile;

blade closely appressed to stem, lanceolate to ovate-oblong, 1.4–5.5 × 0.4–1.2 cm, base attenuate, margins entire, apex acute to rounded, surfaces glabrous or densely erect-hirsute;

bracts usually very reduced, sublinear.

Inflorescences

usually congested, leafy spikes or racemes, flowers solitary in distal leaf axils;

bracteoles attached 1–2.2 mm distal to base of ovary, linear-lanceolate or narrowly elliptic, 3–6.5(–7.2)× 0.3–1.5(–1.7) mm, apex acuminate, surfaces hirtellous.

leafy racemes, flowers solitary in leaf axils;

bracteoles attached in subopposite pairs just proximal to ovary base, lanceolate-linear, 2.5–7 × 0.5–1.5 mm, margins entire, apex acute, surfaces hirtellous.

Flowers

sepals ascending with reflexed tips, pale green abaxially, creamy white adaxially, often tinged with pink or red, ovate-deltate, 3.5–5.5(–6) × 2–4 mm, margins entire, apex elongate-acuminate to subcuspidate, surfaces densely hirtellous;

petals 0;

filaments yellowish, 1.5–2.5 mm, base dilated, anthers 0.6–0.9(–1.3) × 0.5–0.7 mm;

pollen shed in tetrads;

ovary obovoid to cup-shaped, 2.5–4 × 2.5–4 mm;

nectary disc elevated 0.3–0.7 mm on ovary apex, bright yellow, turning black upon drying, 2–3.6 mm diam., indistinctly 4-lobed, densely hirtellous around style base and between lobes;

style cream, 1–2 mm, sparsely to densely hirtellous, especially proximally, stigma capitate, 0.3–0.6 × 0.3–0.6 mm, not exserted beyond anthers.

sepals narrowly ovate-deltate, (5.4–)6–12 × 2.2–4.5(–5) mm, margins entire, apex acute, surfaces hirtellous or, sometimes, glabrous;

petals cordate, 12–14 × 13–14 mm, base attenuate, apex emarginate;

filaments opaque white, awl-shaped, 2–5.5 mm, anthers 1.4–2.6 × 0.4–0.8 mm;

ovary subcuboid or globose, 3–4 × 3–4 mm;

nectary disc elevated, domed, 1–1.4 mm diam., prominently 4-lobed, ringed with stiff, spreading hairs or glabrous;

style 1.3–4.5(–5.5) mm, glabrous, stigma capitate to hemispherical, 0.9–1.5 × 0.9–2(–2.5) mm, often shallowly 4-lobed, as long as stamens, sometimes exserted beyond them.

Capsules

subglobose or, sometimes, oblong-obovoid, subterete or with 4 rounded corners, 3–5 × 3–4.5 mm, hard-walled, irregularly dehiscent, pedicel 0–1 mm.

subcuboid to squarish globose, 4–6 × 4–5 mm, 4-angled, often also 4-winged, wings 0.3–1.1 mm wide, pedicel 3–10 mm.

Seeds

brown, elliptic-oblong or oblong-ovoid, slightly curved on both ends, 0.5–0.7 × 0.3–0.4 mm, surface cells ± isodiametric.

light brown, oblong to reniform, 0.6–0.7 × 0.3–0.4 mm, surface cells elongate transversely to seed length.

2n

= 32.

= 16.

Ludwigia pilosa

Ludwigia hirtella

Phenology Flowering Jun–Sep. Flowering summer–early fall.
Habitat Roadside ditches, marshes, swales in sandy pine flats, edges of pocosins, peaty bogs, low grassy savannas, swamp forests. Boggy depressions, roadside ditches, margins of boggy streams.
Elevation 0–300 m. (0–1000 ft.) 0–300 m. (0–1000 ft.)
Distribution
from FNA
AL; FL; GA; LA; MS; NC; SC; TX; VA
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; DC; DE; FL; GA; KY; LA; MD; MS; NC; NJ; OK; SC; TN; TX; VA
[WildflowerSearch map]
[BONAP county map]
Discussion

The distribution of Ludwigia pilosa is nearly continuous along the Atlantic and Gulf coastal plains, from extreme southeastern Virginia to northern Florida, and west to Louisiana and southeastern Texas. Disjunct populations occur in northern Alabama and central North Carolina.

Ludwigia pilosa is easily distinguished from most others in sect. Isnardia by being densely hirtellous throughout. Its showy sepals and nectary disc attract multiple insect visitors including ants, bumblebees, honeybees, moths, and wasps (C. I. Peng 1989).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

A name often found in synonymy with Ludwigia hirtella is a later homonym: L. hirsuta Pursh 1813, not Desrousseaux 1789.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Ludwigioideae > Ludwigia > sect. Isnardia Onagraceae > subfam. Ludwigioideae > Ludwigia > sect. Ludwigia
Sibling taxa
L. alata, L. alternifolia, L. arcuata, L. bonariensis, L. brevipes, L. curtissii, L. decurrens, L. erecta, L. glandulosa, L. grandiflora, L. hexapetala, L. hirtella, L. lanceolata, L. leptocarpa, L. linearis, L. linifolia, L. maritima, L. microcarpa, L. octovalvis, L. palustris, L. peploides, L. peruviana, L. polycarpa, L. ravenii, L. repens, L. simpsonii, L. spathulata, L. sphaerocarpa, L. suffruticosa, L. virgata
L. alata, L. alternifolia, L. arcuata, L. bonariensis, L. brevipes, L. curtissii, L. decurrens, L. erecta, L. glandulosa, L. grandiflora, L. hexapetala, L. lanceolata, L. leptocarpa, L. linearis, L. linifolia, L. maritima, L. microcarpa, L. octovalvis, L. palustris, L. peploides, L. peruviana, L. pilosa, L. polycarpa, L. ravenii, L. repens, L. simpsonii, L. spathulata, L. sphaerocarpa, L. suffruticosa, L. virgata
Synonyms Isnardia mollis, I. pilosa, L. hirsuta, L. mollis, L. rudis Isnardia hirsuta var. permollis, I. hirtella, L. permollis
Name authority Walter: Fl. Carol., 89. (1788) Rafinesque: Med. Repos., hexade 2, 5: 358. (1808)
Web links